Such problems as that which is presented by the flea’s jumping powers,
though essentially physiological in their nature, have their interest
for us here: because a steady, progressive diminution of activity with
increasing size would tend to set limits to the possible growth in
magnitude of an animal just as surely as those factors which tend to
break and crush the living fabric under its own weight. In the case
of a leap, we have to do rather with a sudden impulse than with a
continued strain, and this impulse should be measured in terms of the
velocity imparted. The velocity is proportional to the impulse (_x_),
and inversely proportional to the mass (_M_) moved: _V_ = _x_/_M_. But,
according to what we still speak of as “Borelli’s law,” the impulse
(i.e. the work of the impulse) is proportional to the volume of the
muscle by which it is produced[47], that is to say (in similarly
constructed animals) to the mass of the whole body; for the impulse is
proportional on the one hand to the cross-section of the muscle, and
on the other to the distance through which it contracts. It follows at
once from this that the velocity is constant, whatever be the size of
the animals: in other words, that all animals, provided always that
they are similarly fashioned, with their various levers etc., in like
proportion, ought to jump, not to the same relative, but to the same
actual height[48]. According to this, then, the flea is not a better,
but rather a worse jumper than a horse or a man. As a matter of fact,
Borelli is careful to point out that in the act of leaping the impulse
is not actually instantaneous, as in the blow of a hammer, but takes
some little time, during which the levers are being extended by which
the centre of gravity of the animal is being propelled forwards; and
this interval of time will be longer in the case of the longer levers
of the larger animal. To some extent, then, this principle acts as a
corrective to the more general one, {30} and tends to leave a certain
balance of advantage, in regard to leaping power, on the side of the
larger animal[49].
But on the other hand, the question of strength of materials comes in
once more, and the factors of stress and strain and bending moment make
it, so to speak, more and more difficult for nature to endow the larger
animal with the length of lever with which she has provided the flea or
the grasshopper.
To Kirby and Spence it seemed that “This wonderful strength of insects
is doubtless the result of something peculiar in the structure and
arrangement of their muscles, and principally their extraordinary power
of contraction.” This hypothesis, which is so easily seen, on physical
grounds, to be unnecessary, has been amply disproved in a series of
excellent papers by F. Plateau[50].
Public-domain text, read in full here on John Shaqi.
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