The measurements taken were those of the girth of the tree, in mm.,
at three feet from the ground. The evergreens included species of
Pinus, Eucalyptus and Acacia; the deciduous trees included Quercus,
Populus, Robinia and Melia. I have merely taken mean values for these
two groups, and expressed the monthly values as percentages of the mean
annual increase. The result (as shewn by Fig. 31) is very much what we
might have expected. The growth of the deciduous trees is completely
arrested in winter-time, and the arrest is all but complete over {120}
a considerable period of time; moreover, during the warm season, the
monthly values are regularly graded (approximately in a sine-curve)
with a clear maximum (in the southern hemisphere) about the month of
December. In the evergreen trees, on the other hand, the amplitude
of the periodic wave is very much less; there is a notable amount of
growth all the year round, and, while there is a marked diminution in
rate during the coldest months, there is a tendency towards equality
over a considerable part of the warmer season. It is probable that some
of the species examined, and especially the pines, were definitely
retarded in growth, either by a temperature above their optimum, or by
deficiency of moisture, during the hottest period of the year; with
the result that the seasonal curve in our diagram has (as it were) its
region of maximum cut off.
[Illustration: Fig. 31. Periodic annual fluctuation in rate of growth
of trees (in the southern hemisphere).]
In the case of trees, the seasonal periodicity of growth is so well
marked that we are entitled to make use of the phenomenon in a converse
way, and to draw deductions as to variations in {121} climate during
past years from the record of varying rates of growth which the tree,
by the thickness of its annual rings, has preserved for us. Mr. A. E.
Douglass, of the University of Arizona, has made a careful study of
this question[151], and I have received (through Professor H. H. Turner
of Oxford) some measurements of the average width of the successive
annual rings in “yellow pine,” 500 years old, from Arizona, in which
trees the annual rings are very clearly distinguished. From the year
1391 to 1518, the mean of two trees was used; from 1519 to 1912, the
mean of five; and the means of these, and sometimes of larger numbers,
were found to be very concordant. A correction was applied by drawing
a long, nearly straight line through the curve for the whole period,
which line was assumed to represent the slowly diminishing mean width
of ring accompanying the increase of size, or age, of the tree; and the
actual growth as measured was equated with this diminishing mean. The
figures used give, accordingly, the ratio of the actual growth in each
year to the mean growth corresponding to the age or magnitude of the
tree at that epoch.
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