It was at once manifest that the rate of growth so determined shewed a
tendency to fluctuate in a long period of between 100 and 200 years. I
then smoothed in groups of 100 (according to Gauss’s method) the yearly
values, so that each number thus found represented the mean annual
increase during a century: that is to say, the value ascribed to the
year 1500 represented the _average annual growth_ during the whole
period between 1450 and 1550, and so on. These values give us a curve
of beautiful and surprising smoothness, from which we seem compelled
to draw the direct conclusion that the climate of Arizona, during the
last 500 years, has fluctuated with a regular periodicity of almost
precisely 150 years. Here again we should be left in doubt (so far
as these {123} observations go) whether the essential factor be a
fluctuation of temperature or an alternation of moisture and aridity;
but the character of the Arizona climate, and the known facts of recent
years, encourage the belief that the latter is the more direct and more
important factor.
[Illustration: Fig. 32. Long-period fluctuation in rate of growth of
Arizona trees (smoothed in 100-year periods), from A.D. 1390–1490 to
A.D. 1810–1910.]
It has been often remarked that our common European trees, such for
instance as the elm or the cherry, tend to have larger leaves the
further north we go; but in this case the phenomenon is to be ascribed
rather to the longer hours of daylight than to any difference of
temperature[152]. The point is a physiological one, and consequently of
little importance to us here[153]; the main point for the morphologist
is the very simple one that physical or climatic conditions have
greatly influenced the rate of growth. The case is analogous to the
direct influence of temperature in modifying the colouration of
organisms, such as certain butterflies. Now if temperature affects the
rate of growth in strict uniformity, alike in all directions and in all
parts or organs, its direct effect must be limited to the production
of local races or varieties differing from one another in actual
magnitude, as the Siberian goldfinch or bullfinch, for instance, differ
from our own. But if there be even ever so little of a discriminating
action in the enhancement of growth by temperature, such that it
accelerates the growth of one tissue or one organ more than another,
then it is evident that it must at once lead to an actual difference of
racial, or even “specific” form.
It is not to be doubted that the various factors of climate have
some such discriminating influence. The leaves of our northern trees
may themselves be an instance of it; and we have, {124} probably, a
still better instance of it in the case of Alpine plants[154], whose
general habit is dwarfed, though their floral organs suffer little or
no reduction. The subject, however, has been little investigated, and
great as its theoretic importance would be to us, we must meanwhile
leave it alone.
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