[Illustration: FIG. 29.—Head of fœtus, of about 5 weeks, from ventral
aspect (after His), showing the primitive stomodæum bounded above by the
undivided fronto-nasal process, laterally by the maxillary, and below by
the still separate mandibular processes. The quinqueradiate appearance is
well represented. (_Sutton._)]
[Illustration: FIG. 30.—Head of fœtus from ventral aspect of a little
later date (6-7 weeks). The mandibular processes have now united; the
orbito-nasal fissure has come in contact with the ocular vesicle, and the
fronto-nasal process has developed into external and internal nasal (or
globular) processes around the nasal fossæ (after His). (_Sutton._)]
At the same time that this mandibular arch is being developed, other
changes are occurring around the upper part of the stomodæum, viz. the
shutting off of sacs lined with epiblast to assist in the formation of
the organs of special sense, and the outgrowth between them of fleshy
processes which by their later amalgamation form the facial skeleton
and coverings. Three of these involutions of epiblast occur, two
communicating more or less with the stomodæum, viz. the nasal and ocular;
whilst the third, or auditory, is separate. Expansions from the cerebral
vesicles meet them, and by further changes, unnecessary to particularize
here, the organs of special sense are elaborated.
[Illustration: FIG. 31A.—Head of fœtus at a somewhat later date (8 weeks)
as seen from the front, showing the nasal and maxillary processes in
close apposition, and the clefts between them diminishing in size.
FIG. 31B.—The same, seen from below, the mandibular process having been
removed.
_i.m._ Central portion of fronto-nasal process, _m.n.pr._ Internal nasal
process. _pr.gl._ Globular process, or the lower rounded extremity of
the former. _l.n.pr._ External nasal process. _m.x._ Maxillary process.
_m.n._ Mandibular process.]
The most anterior of these depressions are the primary _Olfactory pits_
which appear on either side on the lower surface of the wall of the
anterior cerebral vesicle at a very early date. They are at first
merely depressions surrounded by a raised margin; but subsequently they
become pyriform by the extension of the lower end as a groove into the
stomodæum. Each pit and groove is bounded laterally by thickened rounded
outgrowths developed from a broad median fleshy protrusion from the same
part of the cerebral vesicle, the fronto-nasal process; these are termed
respectively the external and internal nasal processes, the latter being
also named by His the globular processes (Fig. 30).
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