The accessory tooth in the os incisivum met with in some cases of
alveolar harelip is maintained by Albrecht to be a reappearance of this
lost middle incisor; and his explanation of such an occurrence seems very
feasible, viz. that the existence of the alveolar cleft prevents the
naso-palatine artery from anastomosing with the posterior palatine, and
thus the vascular supply to the os incisivum is greater than it should
be under normal circumstances; hence, there being a superabundance of
nutrient material, nature uses such in the restoration of a structural
unit ordinarily suppressed. The same fact (viz. the absence of the usual
anastomosis) may explain why the mesognathion is (even when demonstrably
present) always small and the lateral incisor not infrequently stunted or
absent, and so answers the objection to this theory which has been raised
on the ground that in cases of alveolar harelip an incisor external to
the cleft is not invariably present.
Hence the dental formula in cases of alveolar harelip will vary as
follows:
In tetraprodontous jaws—
CI₂=I₁I₁=I₂C,
or
C=I₁I₁=C;
in hexaprodontous jaws—
CI₂=IₐI₁I₁Iₐ=I₂C,
or
C=IₐI₁I₁Iₐ=C,
where Iₐ represents the accessory incisor, the other letters as on p. 51.
The former of each of these pairs of formulæ represent the mesognathion
and lateral incisor as present, the latter as absent.[59]
Notwithstanding the mass of positive evidence which is steadily
accumulating in favour of Albrecht’s theory, there are still some careful
observers who will not admit its truth. Some attack it on the ground that
the intermaxilla does not consist of four portions (Kölliker, &c.); with
such we have already dealt. Others object to it on the ground that any
teeth existing external to the cleft are merely accessory teeth, or due
to a bifidity of the canine. The arguments with which they support their
opinion are derived from the following considerations:
(i) That in normal development accessory teeth do certainly occur,
as in the case mentioned above (p. 53), whilst in a few instances of
abnormality the same condition has been noted (_e. g._ a few cases
recorded by Kölliker;[60] see also Fig. 19, showing a facial cleft in
which several accessory teeth are present along the inner margin). And
the explanation given of such facts is that the involution of mucous
membrane from which the teeth are developed is continuous along the
alveolar ridge, and not localised to the definite spots from which the
teeth subsequently erupt. This, however, is merely an opinion still _sub
judice_, and not absolutely proven.
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