On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
But even the conjectural explanation now given fails us in the other
cases of local modification. Why the species of the western islands
should be smaller than those further east,—why those of Amboyna should
exceed in size those of Gilolo and New Guinea—why the tailed species of
India should begin to lose that appendage in the islands, and retain no
trace of it on the borders of the Pacific, are questions which we cannot
at present attempt to answer. That they depend, however, on some general
principle is certain, because analogous facts have been observed in
other parts of the world. Mr. Bates informs me that, in three distinct
groups, Papilios which on the Upper Amazon and in most other parts of
South America have spotless upper wings obtain pale or white spots at
Para and on the Lower Amazon; and also that the _Æneas_-group of
Papilios never have tails in the equatorial regions and the Amazons
valley, but gradually acquire tails in many cases as they range towards
the northern or southern tropic. Even in Europe we have somewhat similar
facts; for the species and varieties of butterflies peculiar to the
island of Sardinia are generally smaller and more deeply coloured than
those of the mainland, and _Papilio Hospiton_ has lost the tail, which
is a prominent feature of the closely allied _P. Machaon_.
Facts of a similar nature to those now brought forward would no doubt be
found to occur in other groups of insects, were local faunas carefully
studied in relation to those of the surrounding countries; and they seem
to indicate that climate and other physical causes have, in some cases,
a very powerful effect in modifying specific form, and thus directly aid
in producing the endless variety of nature.
I may state that I can adduce facts perfectly analogous to these from
other families of Lepidoptera, especially the Danaidæ; but as the
greater part of the species are still undescribed, I can only now assert
that similar phenomena do occur there.
_Mimicry._
I need scarcely say that I entirely agree with Mr. Bates’s explanation
of the causes which have led to one group of insects mimicking another
(Trans. Linn. Soc. vol. xxiii. p. 495). I have, therefore, only now to
adduce such illustrations of this curious phenomenon as are furnished by
the Eastern Papilionidæ, and to show their hearing upon the phenomena of
variation already mentioned. As in America, so in the Old World, species
of Danaidæ are the objects which the other families most often imitate.
But, besides these, some genera of Morphidæ and one section of the genus
_Papilio_ are also less frequently copied. Many species of _Papilio_
mimic other species of these three groups so closely that they are
undistinguishable when on the wing; and in every case the pairs which
resemble each other inhabit the same locality.
Public-domain text, read in full here on John Shaqi.
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