On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region — John Shaqi
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
The terms “_mimicry_” and “_mimickers_” have been objected to on the
ground that they imply _voluntary action_ on the part of the insects.
This appears to me of little importance compared with the advantages
of convenience, flexibility, and expressiveness which they undoubtedly
possess, especially as the whole theory propounded by the originator
of the term in this sense excludes all idea of voluntary action. The
only approximately synonymous words, not implying will, are
_resemblance_, _similarity_, and _likeness_; and it is evident that
none of these can be applied intelligibly under the variety of forms
required, and to which Mr. Bates’s expression so readily lends itself
in the terms _mimic_, _mimickers_, _mimicry_, _mimicked_. Add to this
the inconvenience of changing a term which, from the interest and wide
discussion of the subject, must be already very generally understood,
and I think it will be admitted that nothing would be gained by
altering it, even if a better word were pointed out, which has not yet
been done.
We have therefore fifteen species or marked varieties of _Papilio_ which
so closely resemble species of other groups in their respective
localities, that it is not possible to impute the resemblance to
accident. The first two in the list (_Papilio paradoxa_ and _P. Caunus_)
are so exactly like _Euplœa Midamus_ and _E. Rhadamanthus_ on the wing,
that, although they fly very slowly, I was quite unable to distinguish
them. The first is a very interesting case, because the male and female
differ considerably, and each mimics the corresponding sex of the
_Euplœa_. A new species of _Papilio_ which I discovered in New Guinea
resembles _Danais sobrina_, Bd., from the same country, just as _Papilio
Macareus_ resembles _Danais Aglaia_ in Malacca, and (according to Dr.
Horsfield’s figure) still more closely in Java. The Indian _Papilio
Agestor_ closely imitates _Danais Tytia_, which has quite a different
style of colouring from the preceding; and the extraordinary _Papilio
idæoides_ from the Philippine Islands must, when on the wing, perfectly
resemble the _Hestia Leuconoë_ of the same region, as also does the _P.
Delessertii_, Guér., imitate an undescribed species of _Hestia_ from
Penang. Now in every one of these cases the Papilios are very scarce,
while the Danaidæ which they resemble are exceedingly abundant—most of
them swarming so as to be a positive nuisance to the collecting
entomologist by continually hovering before him when he is in search of
newer and more varied captures. Every garden, every roadside, the
suburbs of every village are full of them, indicating very clearly that
their life is an easy one, and that they are free from persecution by
the foes which keep down the population of less favoured races. This
superabundant population has been shown by Mr. Bates to be a general
characteristic of all American groups and species which are objects of
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