On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region — John Shaqi
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
The next pair, _P. Theseus_, Cr., and _P. Antiphus_, Fab., have been
united as one species both by De Haan and in the British Museum
Catalogues. The ordinary variety of _P. Theseus_ found in Java almost as
nearly resembles _P. Diphilus_, Esp., of the same country. The most
interesting case, however, is the extreme female form of _P. Memnon_
(_P. Achates_, Cr.)[10], which has acquired the general form and
markings of _P. Coon_, an insect which differs from the ordinary male
_P. Memnon_, as much as any two species differ which can be chosen in
this extensive and highly varied genus; and, as if to show that this
resemblance is not accidental, but is the result of law, when in India
we find a species closely allied to _P. Coon_, but with red instead of
yellow spots (_P. Doubledayi_, Wall.), the corresponding variety of _P.
Androgeus_ (_P. Achates_, Cram., 182, A, B,) has acquired exactly the
same peculiarity of having red spots instead of yellow. Lastly, in the
island of Timor, the female of _P. Œnomaus_ (a species allied to _P.
Memnon_) resembles so closely _P. Liris_ (one of the _Polydorus_-group),
that the two, which were often seen flying together, could only be
distinguished by a minute comparison after being captured.
Footnote 10:
See Plate I. fig. 4.
The last six cases of mimicry are especially instructive, because they
seem to indicate one of the processes by which dimorphic forms have been
produced. When, as in these cases, one sex differs much from the other,
and varies greatly itself, it may happen that occasionally individual
variations will occur having a distant resemblance to groups which are
the objects of mimicry, and which it is therefore advantageous to
resemble. Such a variety will have a better chance of preservation; the
individuals possessing it will be multiplied; and their accidental
likeness to the favoured group will be rendered permanent by hereditary
transmission, and, each successive variation which increases the
resemblance being preserved, and all variations departing from the
favoured type having less chance of preservation, there will in time
result those singular cases of two or more isolated and fixed forms
bound together by that intimate relationship which constitutes them the
sexes of a single species. The reason why the females are more subject
to this kind of modification than the males is, probably, that their
slower flight, when laden with eggs, and their exposure to attack while
in the act of depositing their eggs upon leaves, render it especially
advantageous for them to have some additional protection. This they at
once obtain by acquiring a resemblance to other species which, from
whatever cause, enjoy a comparative immunity from persecution.
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