On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
_Hab._ Malacca, Sumatra, Borneo (♂, ♀) (_Wall._), Assam (_Brit.
Mus._).
h. _Pammon_ group.
62. PAPILIO PAMMON, Linnæus. Tab. II. figs. 1 (♂), 3, 5, 6 (♀ ♀).
♂, _P. Pammon_, L.; Cram. Pap. Ex. t. 141. f. B; Boisd. Sp. Gén. Lép.
p. 272.
♀, _P. Polytes_, L.; Cram. Pap. Ex. t. 265. f. A, B, C.
_Hab._ Malacca, Singapore (_Wall._), China, India, Ceylon.
The continental specimens of _P. Pammon_ have all considerably developed
tails in both sexes; the insular specimens on the other hand, (which I
treat as a separate species), have only a prominent tooth or very short
tail in the males. The females also differ considerably, presenting an
analogous but distinct series of forms. In the true _P. Pammon_ the
males are very constant; but the females exist under three distinct
forms, each of them presenting more or less numerous varieties, viz.:—
_1st form_ of female. Tab. II. fig. 3.
This exactly resembles the male, except in the possession of a distinct
ocellus at the anal angle on the upper surface. Rarely a variety occurs
having in addition a submarginal row of red lunules, indicating a slight
approximation towards some varieties of the second form.
_2nd form_ of female (_P. Polytes_). Tab. II. fig. 5.
This is by far the most common form of female. A variety of this rarely
occurs, which wants the red patch at the anal angle, and has the white
patch formed of a row of spots all situated a little below the discoidal
cell. This is the nearest approach to the first form.
_3rd form_ of female (_P. Romulus_, Cram. Pap. Ex. t. 43. f. A; _P.
Mutius_, Fab., Boisd. Sp. Gén. p. 270; _P. Hector_ ♀, De Haan). Tab. II.
fig. 6.
This not uncommon Indian butterfly I consider to be a third form of the
female of _P. Pammon_. I was first led to suspect this by finding that
no males of it are known (the male and female from Ceylon, noted in the
British Museum List, I have ascertained to be both females), nor have I
been able to find any after an examination of the chief collections in
England. It is also to be observed that it has been received from no
locality which is not also inhabited by _P. Pammon_; there is no other
known Indian butterfly that can possibly be the other sex of it; and
lastly, it agrees very closely with the second form of female (_P.
Polytes_) in all its details of form, texture, and neuration; and though
at first sight having a very different aspect, specimens are to be found
which by a very slight modification could be changed so as to resemble
that form. I am therefore quite satisfied in my own mind that I am right
in sinking this species into a form of _P. Pammon_. I have already
stated my opinion that it mimics _P. Hector_, with which, however, it
has no affinity. The resemblance was such as to induce De Haan to place
it as the female of that species.
63. PAPILIO THESEUS, Cramer. Tab. II. figs. 2, 4, 7 (♀ ♀).
_P. Theseus_, Cr. Pap. Ex. t. 180. f. B (♀); Boisd. Sp. Gén. Lép. p.
276.
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