On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of GeneraWollaston, Thomas Vernon
Science
On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of Genera
Wollaston, Thomas Vernon
Insects; Variation (Biology)
In exact conformity with what the above remarks will have prepared us
for, we find that the _Dromius obscuroguttatus_, Dufts., of Central
Europe, has undergone on the mountain summits of Madeira changes
precisely to that extent which we should have calculated upon; and
although they would seem in reality to be referable to climate _and
isolation_ combined, yet, since it is not always possible (as lately
stated) to treat the elements of disturbance separately, and it is my
object in this short treatise to bring forward a few prominent
examples in support of the considerations proposed, rather than to
accumulate a mass of material for the registry of which my space would
be inadequate, I will quote _in extenso_ the reflections which, during
the compilation of the 'Insecta Maderensia,' suggested themselves to
me. "The _Dromius obscuroguttatus_ is a common European insect, and
the Madeiran specimens recede from the ordinary ones in being slightly
larger, and in having their elytra more obscurely striated, with the
humeral patch less distinct: their entire surface, moreover, is of a
deeper black, a difference which is especially perceptible on the
legs. It occurs in the greatest profusion in Madeira proper, though
only from about 5000 to 6000 feet above the sea. Although so common
throughout Europe, it is perhaps, when geographically considered, one
of the most interesting of the Madeiran Coleoptera, as affording a
striking example, not only of the modification of form in a normally
northern insect when on its southern limit, but as showing likewise
how a species, abundant on the low sandy shores and sheltered
sea-cliffs of more temperate regions, finds its position here only on
the summits of the loftiest mountains. It is true that the aberration
from the typical state is not in the present instance very
considerable; yet when the circumstances producing it are taken into
account, I am persuaded that the difference is exactly of that nature
on which too great stress cannot possibly be placed, when discussing
the general question of geographical distribution as having a
tendency, more or less directly, to affect both colour and form. It is
well known to naturalists that a multitude of insects from the New
World, receding from their European analogues merely in certain
excessively minute characters, have usually been pronounced at once as
new to science, first because those differences are constant, and
secondly because the specimens have been received from the other side
of the Atlantic. And yet in instances like the present one,--in an
island which, while it belongs artificially to Europe, is yet
naturally sufficiently distinct from it as to form at any rate a
stepping-stone to the coast of Africa and the mountains of
Barbary,--species similarly circumstanced are not necessarily received
as new (and rightly so, I apprehend), though in every respect
affording differences not only _analogous_ to those already mentioned,
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