On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of GeneraWollaston, Thomas Vernon
Science
On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of Genera
Wollaston, Thomas Vernon
Insects; Variation (Biology)
And if we inquire, on the other hand, into the _aboriginal_ species of
those islands,--or, at any rate, into such of them whose naturally
acquired range embraces the opposite extremes of atmosphere,--we shall
detect no less surely (albeit within a narrower space) the result of
climatal action on insect form. The _Helops confertus_, Woll., "varies
according to the altitude at which it is found; being usually deeply
striated and rugose on its lower, but subpicescent and much more
lightly sculptured on its upper limits. I have taken specimens indeed
on Pico Ruivo, and on the mountain-plain of the Fateiras, which are so
far diminished in roughness as almost to resemble, at first sight, the
_H. Pluto_[11]." The _Pecteropus Maderensis_, Woll., which ranges from
about 2500 feet above the sea to the summits of the loftiest hills,
although usually with pale legs, is distinguished by having its femora
almost invariably dusky when on its highest elevation; and, following
out the analogy with that beetle, the _Trechus alticola_, Woll.,
should perhaps be regarded as an alpine state of the _T. custos_. The
_Calathus complanatus_, Koll., assumes along the upland heights a very
different aspect to what it does in the regions below, being generally
more piceous and convex, altogether broader (in proportion) and
shorter, and with _both_ sexes (though, of course, especially the
male) shining.
Nor is this principle of topographical variability (the result of
climate) less apparent in other countries also. The _Notiophili_, for
instance, "are extremely unstable, both in their sculpture and hue,
being subject to considerable local modifications, though more
particularly affected, it would appear, by altitude. Thus, in our own
country, the _N. semipunctatus_, Fab., one of the common
representatives of the plains, is found likewise on the summits of the
mountains; but at that elevation it becomes liable to great
alternations of colour, ranging from pale brassy-brown, with the apex
testaceous, into deep black. The sculpture, however, perhaps is nearly
as much dependent on other circumstances for its modification as upon
altitude, since it seems tolerably clear that proximity to the
sea-shore, especially where the localities are saline, will frequently
produce a more faintly impressed surface[12]." It has indeed been
lately suggested, that the _Helobia nivalis_, Payk., may be perhaps,
after all, but a mountain variety of the _H. brevicollis_; the
_Leistus montanus_, Steph., of the _L. fulvibarbis_, and the _Patrobus
septentrionis_, Dej., of the _P. excavatus_; but of this I think
further proof is needed, seeing that certain species do appear to
exist which are _strictly_ alpine (that is to say, which have not
been, severally, detected in the lower regions of more northern
zones); and, in _most_ instances, where aberrations are to be met
with from the effect of _altitude_, we have a right to inquire
(provided the types from which they are supposed to have originally
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