On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of GeneraWollaston, Thomas Vernon
Science
On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of Genera
Wollaston, Thomas Vernon
Insects; Variation (Biology)
In the Madeiras, this diminution and enlargement of stature,
accompanied for the most part respectively by the retention and
annihilation of the powers of flight, is singularly traceable on the
selfsame rocks, particularly the smaller ones of the group. Thus, on
the Flat Deserta, or Ilheo Chao, the _Scarites abbreviatus_, Koll.,
_Laparocerus morio_, Schoen., and the _Helops Vulcanus_, Woll., attain
a gigantic size; yet it is on that very island that the _Ptinus
albopictus_, Woll., finds its minimum of development,--scarcely
exceeding in dimensions some of the larger members of the
_Trichopterygia_. The Deserta Grande has some special modifying
capability of its own,--the _Eurygnathus Latreillei_, Lap.,
_Notiophilus geminatus_, Dej., _Zargus pellucidus_, Woll., _Calathus
complanatus_, Koll., _Olisthopus Maderensis_, Woll., _Caulotrupis
conicollis_, Woll., _Laparocerus morio_, Schoen., _Omias Waterhousei_,
Woll., _Helops Vulcanus_, Woll., and the _Ellipsodes glabratus_, Fab.,
being also larger on that rock than is typical: all of them, however,
with the exception of _Notiophilus geminatus_, are there, as
elsewhere, apterous.
Other qualifying results, from isolation, are equally apparent. Take
_colour_, for instance; and we shall perceive that in the _Dromius
sigma_, Rossi, it is sensibly affected. The normal state of that
insect "does not occur at all in Madeira proper, but only in Porto
Santo. True it is that the modifications in the several islands
present but slight differences _inter se_; nevertheless, being
constant, I would lay particular stress upon them, since they go very
materially to prove that the effects of isolation on external insect
form are even more important, if possible, than those of latitude.
That this is the case in the present instance, appears clear from
facts so minute as these. For, out of the many specimens which have
come under my observation from various countries of Europe, if there
is one point more constant than another in this otherwise variable
species, it is, I believe, to all circumstances, its immaculate
prothorax. Now, whilst this (we may almost say essential) character
obtains in Porto Santo, in Madeira it does not hold good: the
prothorax there is invariably infuscate in the centre; and on a small
adjacent rock (the Ilheo de Fora) it is entirely dark. Nor let anyone
suppose that details apparently so trivial are beneath our notice, or
the mere result of chance, since it is by the observation of such-like
points, and by marking their development according to the
circumstances of the several localities in which they obtain, that we
are alone able to appreciate their importance, and so to form, in a
wider and geographical sense, a correct estimate of their value[40]."
The _Olisthopus Maderensis_, Woll., is much paler, larger, and more
opake, on the Dezerta Grande than it is in Madeira proper. So great
indeed is the change which it has undergone through a long isolation
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