On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of GeneraWollaston, Thomas Vernon
Science
On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of Genera
Wollaston, Thomas Vernon
Insects; Variation (Biology)
on that rock, "that, had the case been a solitary one, I should not
have hesitated in regarding the specimens obtained from thence as
specifically distinct; nevertheless, with the knowledge both of the
modifying effects of isolation, and also of the _kind_ of modification
essentially peculiar to that island, I am perfectly satisfied that it
is a mere local state, although a very remarkable one, and has no
claim whatsoever to be otherwise considered[41]." The _Pecteropus
Maderensis_, Woll., is of a greenish-brassy tinge in Porto Santo, and
much acuminated in front; whereas on the Dezerta Grande it is almost
invariably _coppery_, and less narrowed anteriorly. The _Caulotrupis
lucifugus_, Woll., although ranging through no very opposite phases,
either of outline or sculpture, "appears to possess a slight
modification for every island of the Madeiran Group: and hence small
shades of difference, which might otherwise be regarded as trifling,
become directly important, and cannot be ignored in a local
fauna,--even though a general collector may deem it unnecessary to
recognize them. In real fact, however, such distinctions, when viewed
geographically, are of the greatest interest, as serving to illustrate
what we have so often had occasion to comment upon, namely the
influence of isolation and other circumstances on external insect
form[42]." The _Psylliodes vehemens_, Woll., is permanently paler in
Porto Santo than it is in Madeira proper, being almost entirely
testaceous. "That the species is identical, however, with the Madeiran
one I have not the slightest doubt,--the sculpture and colour, as I
conceive, having merely undergone a change since the remote period
of its isolation on a comparatively calcareous soil[43]." The
_Scarites abbreviatus_, Koll., occupies the loftiest peaks of nearly
all the Madeiran islands, and was probably once abundant over the
entire ancient continent, whatsoever its limits may have been, of
which the present group forms but an isolated part. "There are traces
of it in the Canaries, from whence occasional specimens have been
brought, and which, from the want of local data and of sufficient
numbers to reason upon, have in their turn been severally regarded as
distinct. The fact however is, that the species in question is an
extremely variable one, assuming differences of size according to the
altitude at which it lives, and differences of sculpture according to
the circumstances of the spot on which it is isolated. That such is
actually the case, a careful observation of the many minute changes
which the insect has undergone in the various islands and altitudes of
the Madeiran group will, I think, prove to a demonstration. For it is
impossible to suppose that every rock contains its own _species_, that
is to say, has had a separate creation expressly for itself,--a
conclusion at which we must assuredly arrive, if small and even
constant differences are _of necessity_ specific. Rejecting therefore
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