On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of GeneraWollaston, Thomas Vernon
Science
On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of Genera
Wollaston, Thomas Vernon
Insects; Variation (Biology)
Such are a few of the effects which elevation and subsidence, even on
a small scale, would seem (when tested by theory and practice) to
produce. It yet remains for us to suggest, that the latter, when
carried to its maximum, so as to cause the actual separation by the
sea of one district from another, is a contingency of immense
significance in regulating the distribution of the Annulose tribes.
Their outward contour and aspect we have shown in a previous chapter
to be very largely beneath the control of isolation, provided a
sufficient _time_ can be granted for the change: but their ultimate
absence from any particular place, through the impediment which it
offers to their migratory progress, we have not yet touched upon. Let
us conceive, therefore, an extensive continent; and, since the
insects which at present inhabit our earth must, if the doctrine of
specific centres be true, have been originally created in certain
definite spots, let us suppose a limited proportion of them to have
been first produced upon this tract. Self-dissemination, we will
assume, has been going on for centuries: those species which were
gifted with quick diffusive powers have become pretty evenly dispersed
over its surface; whilst those of naturally slow or sedentary habits
have peopled, comparatively, but small areas around the respective
localities of their birth. Such may have been the case, at some fixed
period, amongst the aboriginal beings of any country which we choose
to select as an illustration. But there is another element to be
considered. If this region be not insular, it will have received
colonists from foci of radiation situated beyond its bounds; and
these, therefore, according to their several capabilities for
progression, will have, likewise, in parts, overspread, or tenanted,
it. Now it is impossible to cite a more simple example than this. But
let us endeavour to realize what would be the necessary consequence of
the breaking up of such a district as that which we have imagined. If
a _general_ sinking should take place, causing its higher points to be
alone visible above the ocean, or merely a _partial_ one, so as to
admit of the sea encompassing portions of it which would remain
unaffected in their altitude; the result practically would be the
same,--namely, the constitution of a group of islands out of a once
continuous land. Then, as regards the animal population of this
tract, the main phaenomena are almost self-evident. Should any of its
isolated fragments chance to contain a portion of one of _those
limited areas_ which a species of slow progressive powers had
succeeded in colonizing, it would of course harbour (provided that the
other portion has disappeared) what would now be defined as _endemic_.
Numbers of these small areas, or, in other words, of the species which
had overspread them, would in all probability be lost for ever; whilst
the occurrence of any of the surviving ones in more than a single
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