On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of GeneraWollaston, Thomas Vernon
Science
On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of Genera
Wollaston, Thomas Vernon
Insects; Variation (Biology)
island would manifestly depend on the proximity of the islands _inter
se_. Those forms which had diffused themselves over the whole original
continent would now be found in all the detachments of the cluster;
whilst others, which had wandered over the greater portion of it only,
might be traceable perhaps in every island _except a few_.
Such are the primary facts which suggest themselves, whilst discussing
the question of isolation as regulating the _distribution_ of the
Annulose tribes. Its _after effects_, on their external configuration
and development, we have examined in a preceding chapter of this
treatise; and we have also lately intimated what might be a few of the
presumptive consequences of a subsidence (in a general sense), _apart
from_ the still more important principle of isolation. Before,
however, we dismiss these brief and elementary reflexions on the
upward and downward movements which geology testifies to have
occurred, at various epochs, on the earth's surface, I shall perhaps
be pardoned if I digress so far from my immediate subject as to trace
out some of the actual results of isolation in the diffusion of the
Insecta (especially recognizable in the stoppage of a former migratory
progress) in a few of the northern Atlantic groups. I should premise,
however, that it is from the Coleoptera alone that I shall attempt to
draw my inferences; nevertheless, since that order is more extensive
than any of the others, and has moreover been closely investigated in
most of those islands, it may possibly afford us data of sufficient
comprehensiveness and accuracy for practical purposes.
To commence, then, with the Madeiras and Canaries; the first facts
which isolation discloses to us, concerning the statistics of a region
which was once continuous throughout that portion of the Atlantic, are
the _slowness_ and the _direction_ of the ancient migratory movements.
The former of these is rendered evident from the vast number of
endemic species which are at present contained, not merely in the two
groups combined, but in the several islands of which each of them is
composed. True it is, that these peculiar forms are, most of them,
apterous, and of naturally sluggish self-disseminating powers; yet,
still the circumstance remains, that these various creatures had not
overrun areas of any extent before the land of passage was
destroyed,--for otherwise they must have occurred, now, on islands and
rocks but slightly removed from each other, _which they do not_. The
latter of the above conclusions, namely, the _direction_ of the
migratory current, will become apparent in the sequel. We may premise
however, that, so far as the aborigines of this province are
concerned, their course will be found, upon the whole, to have been a
_northerly_ one.
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