The _exoccipitals_ are large, of very great width, but rather low
from above downwards. The position of the foramina which perforate
these bones is peculiar; it resembles more the arrangement seen in
the skull of the alligator than in that of the crocodile, but it has
an additional foramen. There are, then, two small venous foramina
near the condyle; while along the lateral margin of the occiput,
are placed in a vertical line the foramina for the hypoglossal and
pneumogastric nerves, and the internal carotid artery. The foramen
for the facial nerve, etc., is situated in the usual place, and is of
the usual size. The foramen magnum is heart-shaped, low, wide above,
contracting below. The paroccipital processes are long and slender,
and project strongly backwards.
The _supra-occipital_ is very small. It shows to some extent on
the upper surface of the skull, wedged in between the parietals.
As a whole, the occiput is of remarkable shape: it is perfectly
vertical, as in all other crocodilians; is remarkably high from
above downwards, and is very broad at top, becoming extremely narrow
distally. This latter feature is owing to the peculiar shape of the
pterygoids; which, when viewed from behind, do not appear to reach
the basioccipital.
The _basisphenoid_ is so destroyed by crushing, that nothing can be
said of its shape or of its foramina. It was, however, evidently
very stout. The alisphenoid is large and smooth and is not ridged;
the _foramen ovale_ is small, and is not produced into an anterior
notch. The suture between the alisphenoid and the pro-otic are quite
distinct, but the latter is so firmly anchylosed to the quadrate that
its limits are indeterminable. None of the other periotic bones are
visible.
The _quadrate_ is very long and broad. In shape it resembles the
corresponding bone of the alligator, but is somewhat broader. The
lower surface is divided into two unequal parts by a conspicuous
ridge, which runs to within an inch of the articular surface. This
surface has a more decidedly grooved or trochlear appearance than in
the common crocodile; but not to the same degree as in the specimen
described by Dr. Leidy. It is more like the Mississippi alligator in
this respect than any living species of which we have specimens.
Public-domain text, read in full here on John Shaqi.
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