Phylogeny of the Waxwings and Allied BirdsArvey, M. Dale (Martin Dale)
Science
Phylogeny of the Waxwings and Allied Birds
Arvey, M. Dale (Martin Dale)
Bombycillidae -- Phylogeny; Waxwings -- Phylogeny
It may be noted that _Phainoptila_ and _Bombycilla_ have the shortest
arm in the family Bombycillidae. The humerus, radius and ulna are
comparable to the same elements in thrushes and the catbird, and it is
only the extremely short manus in _Phainoptila_ that affects the
total. The manus in _Phainoptila_ is comparatively smaller than in any
other genus of the family Bombycillidae, and this indicates poor
flight power. _Bombycilla_ has a total length corresponding closely to
that in warblers, but the lengths of the distal elements correspond
closely to those in the catbird and thrushes. Of the three segments,
the humerus is, relatively, the most shortened. Next in order of
increasing length of arm is _Dulus_; measurements for it are roughly
the same as those of _Myadestes_. The wing bones of the
Ptilogonatinae, other than _Phainoptila_, are the longest in this
series, and they most nearly resemble the same bones in flycatchers,
Parids, and gnatcatchers.
[Illustration: Fig. 47. Graph showing relative lengths of bones of
the arm. The percentage values are shown on the axis
of the ordinates.
A. _Bombycilla cedrorum_; B. _Bombycilla garrula_;
C. _Dulus dominicus_; D. _Phainoptila melanoxantha_;
E. _Phainopepla nitens_; F. _Ptilogonys cinereus_;
G._ Ptilogonys caudatus_.
a. humerus; b. radius; c. ulna; d. manus; e. total.]
It is notable that, in general, birds with long and narrow wings
appear to have relatively the shortest humeri, with the distal bones,
especially the manus, variable in length and seemingly correlated with
the manner of feather attachment. Those birds with rounded and short
wings have the longest humeri. In swallows, for example, the humerus
is short, whereas the other arm bones are long, and the manus is
unusually large and heavy. A short humerus gives better lever action
in the flight stroke than a long humerus does.
MUSCULATURE
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