The palæotherium was of this period, and also the mastodon, both of
large dimensions. Various families, and even genera, of quadrupeds now
existing were associated with these extraordinary creatures, though of
extinct species, such as the elephant, rhinoceros, hippopotamus, tapir,
horse, bear, wolf, hyæna, weasel, beaver, ox, buffalo, deer, &c.; and
also marine mammalia, as dolphins, sea-calves, walruses, and lamantines.
Indeed, in the constant increase of animal life manifested throughout
the whole of the tertiary strata, the forms approach nearer to living
species as their remains lie high in the series.
In the older Pleiocene period some of the large amphibious quadrupeds,
and other genera of mammalia of the earlier tertiary periods, appear no
more; but there were the mastodon, and the Elephas primigenius or
mammoth, some species of which, of prodigious size, were associated with
numerous quadrupeds of existing genera, but lost species. Extinct
species of almost all the quadrupeds now alive seem to have inhabited
the earth at that time; their bones have been discovered in caverns;
they were embedded in the breccias and in most of the strata of that
epoch—as the hippopotamus, rhinoceros, elephant, horse, bear, wolf,
water-rat, hyæna, and various birds. It is remarkable that in the
caverns of Australia the fossil bones all belong to extinct species of
gigantic kangaroos and wombats, animals belonging to the marsupial
family, which are so peculiarly the inhabitants of that country at the
present day, but of diminished size. The newer Pleiocene strata show
that the same analogy existed between the extinct and recent mammalia of
South America, which, like their living congeners, as far as we know,
belonged to that continent alone; for the fossil remains, quite
different from those in the old world, are of animals of the same
families with the sloths, anteaters, and armadilloes which now inhabit
that country, but of vastly superior size and different species. In
fact, there were giants in the land in those days. Were change of
species possible, one might almost fancy that these countries had
escaped the wreck of time, and that their inhabitants had pined and
dwindled under the change of circumstances. The megatherium and Equus
curvidens, or extinct horse, had so vast a range in America, that, while
Sir Charles Lyell collected their bones in Georgia in 33° N. latitude,
Mr. Darwin brought them from the corresponding latitude in South
America. The Equus curvidens differed as much from the living horse as
the quagga or zebra does, and the European fossil horse is also probably
a distinct and lost species.
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