Physiology: The Science of the BodyMartin, Ernest G.
Science
Physiology: The Science of the Body
Martin, Ernest G.
Physiology
Shortly after food begins to enter the stomach churning movements are
set up. These consist of regular contractions of the smooth muscle
beginning at about the middle of the stomach in the form of a ring and
so causing a deep depression to form right around the stomach, which
then travels along toward the sphincter at the outlet. This, of course,
crowds the food up toward that end; but if the sphincter does not relax,
the food instead of escaping simply squirts back through the space left
in the middle of the ringlike construction and so is actively churned. A
regular procession of these waves travels over this part of the stomach
during the whole time that gastric digestion is going on. The outer half
of the stomach, the side toward the esophagus, does not join in this
churning motion. The walls here remain quiet, merely pressing upon the
food to crowd it up into the part where the churning is going on as fast
as room is made for it there. At intervals during this process the
sphincter relaxes and a small mass of food is crowded through into the
small intestine; then the sphincter closes again, preventing more from
passing until the former mass has had time to be acted upon by the
digestive juices in the intestine. The way in which this sphincter is
controlled is one of the very interesting facts of physiology, but
before describing it we shall have to say something about actual gastric
digestion.
The juice which the glands in the stomach wall secrete is called gastric
juice. This contains three important constituents, first of which is
hydrochloric acid. From the standpoint of chemistry it is a very
interesting thing that the gastric glands should he able to manufacture
a mineral acid like hydrochloric acid. That they do so, however, is
proven by the facts of digestion in everybody. In addition to
hydrochloric acid gastric juice contains two enzymes; the first is
pepsin, which begins the digestion of protein foods, although it does
not carry it through to completion. The other constituent is the enzyme
rennin, which clots milk; this property appears to be useful in that
clotted milk will not pass the sphincter into the small intestine as
quickly as it would if it were liquid, and so the digestive processes
can go on in milk after clotting just as it goes on in any other of our
foods. Since milk is the chief, and in many cases the only, food of the
young it is of course very important that its digestion should be very
efficiently carried on. As the food is churned by the stomach muscles,
it becomes mixed with gastric juice. Any milk that is present will be
clotted and any protein will begin to be digested. The food will also be
mixed with hydrochloric acid. It happens that the salivary enzyme,
ptyalin, will not act in the presence of an acid, so that as soon as any
of the food in the stomach comes in contact with hydrochloric acid of
the gastric juice, the digestion of starch by ptyalin stops. Since
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