The San Josecito collection contains 22 rami of a species of
_Cryptotis_. Many are nearly complete although none possesses the
incisor. In addition there is a rostrum that on the right side bears
the last two unicuspids, P4, M1, and M2. I have compared these fossils
with specimens of the following species of _Cryptotis_: _C. mexicana_,
_C. magna_, _C. nelsoni_, _C. thomasi_, _C. alticola_, _C. parva_, _C.
orophila_, _C. pergracilis_, _C. guerrerensis_, _C. obscura_, _C.
mera_, _C. soricina_, _C. fossor_, _C. goodwini_, _C. griseoventris_,
_C. meridensis_, _C. mayensis_, and _C. micrura_. The four species
first mentioned and the fossils seem to fall into one group. The
remaining species fall into another group characterized by a smaller
occlusal area of the talonid on all three molar teeth with respect to
the trigonid, and especially by the smaller and weaker talonid of m3
which possesses only one bladelike cusp, the hypoconid. In the first
four species the talonids are larger than in the other species when
compared to the trigonids, and the talonid of m3 possesses a well
developed hypoconid and entoconid with a distinct basin between them.
The rami of San Josecito specimens closely resemble those of _C.
mexicana_ in both size and qualitative characters. The rostrum
mentioned above differs from those of _C. mexicana_ in that the
unicuspids are larger, especially the posteriormost one. _Cryptotis
thomasi_ and _C. magna_ are eliminated from consideration here on
geographical grounds. Little difference may be seen between the rami of
_C. mexicana mexicana_ from Veracruz and _C. nelsoni_. The fossils are
referred to the former species since it has a rather wide distribution
in México in contrast to _C. nelsoni_ which is restricted to Volcán de
Tuxtla, Veracruz. The northernmost Recent occurrence of _C. mexicana_
known to me is from Las Vigas, Veracruz. As far as I know the species
has not previously been recorded from the Pleistocene.
Stirton (1930:225), in summarizing the group characters of Recent
Soricidae, listed the bicuspid talonid of m3 as one of the characters
of the "_Blarina_ group," which includes _Neomys_, _Soriculus_,
_Notiosorex_, _Chimarrogale_, _Cryptotis_, and _Blarina_. That this
character does not obtain universally within the group is demonstrated
by the unicuspid structure of the talonid of m3 of the majority of the
species of Mexican _Cryptotis_.
I am grateful to Drs. David E. Johnson and Henry W. Setzer
of the United States National Museum, and to Dr. John
Aldrich and Miss Viola S. Schantz of the United States Fish
and Wildlife Service for permitting me to examine specimens
in their care. Also I am obliged to Professor E. Raymond
Hall for permission to study the specimens from San Josecito
Cave. The late Professor Chester Stock intrusted the
specimens to Professor Hall for study and description.
LITERATURE CITED
CUSHING, J. E., JR.
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