The relations of the two species _Helix nemoralis_ and _hortensis_
provide an illustration of another kind of manifestation of local
peculiarity. _H. hortensis_ and _nemoralis_ as usually met with, are
two very distinct forms. _H. hortensis_ is smaller and duller, and its
peristome is white. _H. nemoralis_ is larger and more shiny, and its
peristome is brown. In several anatomical points, moreover, especially
in the shape of the dart, there are great differences. For a full
account of these peculiarities of the two forms and a discussion of
their inter-relations the reader is referred to the elaborate work of
A. Lang[14] who has studied them extensively and has also succeeded in
experimentally raising hybrids between them. These hybrids were in a
slight degree fertile with both the parent species, but up to the time
of publication no young had been reared from hybrids _inter se_.
Coutagne describes the result of collections made in 62 French
localities. Some had exclusively _hortensis_, some exclusively
_nemoralis_, and in some the two were found in association. He gives
details of five of these collections from which I take the following
summary of the more essential facts, omitting much that is almost
equally significant.
_Locality A_, near Honfleur. Both forms present, each sharply and
normally distinguished, without any intermediates. They are thus found
in many places. Coutagne instances Müller's observations in Denmark, his
own series from the Jura, etc.
_Locality B._ Vonges (Côte d'Or), 242 _hortensis_ taken at random,
showed 128 with light peristomes (either more or less pinkish or
quite white) and 114 with dark _brown_ peristomes; together with 26
_nemoralis_ all with the usual brown peristomes.
Of the _hortensis_ 50 were in ground-colour _opalescens_ and 1 _roseus_;
and in shape 5 were _umbilicatus_.
_Locality C_, about 3 kilometres from _B_. There were found 35
_hortensis_, of which 20 had light peristomes and 15 brown; together
with 7 _nemoralis_.
Of the _hortensis_ none were _opalescens_; 18 were _roseus_ and none has
the shape of _umbilicatus_.
_Locality D_, about 1,200 metres from _B_. 147 _hortensis_, of which 4
had light peristomes and 143 had brown. No _nemoralis_ were found.
None of the _hortensis_ were _opalescens_ or _roseus_, but 30 were
_umbilicatus_.
In these localities intermediates of every grade existed between the
well-characterised _opalescens_, _roseus_, or _umbilicatus_, and the
other forms, but there were no intergrades between the other _nemoralis_
and the smaller _hortensis_, about which there was no hesitation. In the
next locality a very different state of things was found.
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