_Locality E._ Banks of the Yvette at Orsay (Seine-et-Oise). The
actual numbers are not given, but we are told that 58 per cent. were
_hortensis_, 33 per cent. _nemoralis_, and 9 per cent. intermediate. As
at Honfleur, the _hortensis_ had white peristomes, and the _nemoralis_
brown. Coutagne's visits to this locality were in 1878 and 1880, and he
calls attention to the fact that Pascal found similar intermediates in
the same neighbourhood in 1873.
The two species, in Coutagne's view, when they occur together, can
generally be sorted from each other with perfect confidence, and it is
only in exceptional localities that these intermediates occur. Whether
they are hybrids, or whether sometimes the species in their variations
transgress their usual limitations is regarded both by Coutagne and
by Lang as a question not yet answerable with certainty. Coutagne
moreover lays stress on the fact that although each species may be
easily known from the other _in its own district_, yet when shells from
different districts are brought together it is sometimes impossible to
sort them. He mentions an example of such casual intermixture occurring
under natural conditions on an island in the Rhone, to which it may
well be supposed that floods had brought immigrants from miscellaneous
localities. This population contained a very large number of uncertain
specimens, and as he says, it was much as if he were to mix the shells
from his 62 localities, after which it would certainly be impossible to
separate the two species again.[15]
Further evidence is given in the same treatise as to other examples
of polymorphism, especially in the genus _Anodonta_, of which Locard
made 251 species for France alone. Here again are cases like those
already given, and many forms or "modes" are found restricted to special
localities, while occasionally in the same locality dissimilar forms are
found, collectively forming a colony, without intermediates.
Taken as a whole the evidence shows the following conclusions to be
true. Local races, whether of animals or plants, may be distinguished
by characters which we are compelled to regard as trivial, or again by
features of such magnitude that if they were known to us only as the
characteristics of a uniform species they would certainly be assumed
without hesitation to be essential for its maintenance. Local forms
may be sharply differentiated from the corresponding populations of
other localities or they may be connected with them by numbers of
intermediates. Not rarely also we find a fact which has always seemed to
me of special significance, that the peculiarity of the local population
or colony may show itself in a special liability to variation, and
this variability may show itself in one of many degrees, either in the
constant possession of a definite aberration, in a dimorphism, or in an
extreme polymorphism.
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