I am well aware that some very eminent systematists regard the whole
problem as solved. They hold as Darwin did that specific diversity
has no physiological foundation or causation apart from fitness, and
that species are impermanent groups, the delimitations of which are
ultimately determined by environmental exigency or "fitness." The
specific diversity of living things is thus regarded as being something
quite different in nature from the specific diversity of inorganic
substances. In practice those who share these opinions are, as might
be anticipated, to be found among the 'lumpers' rather than among the
'splitters.' In their work, certainly, the Darwinian theory is actually
followed as a guiding principle; unanalysed inter-gradations of all
kinds are accepted as impugning the integrity of species; the underlying
physiological problem is forgotten, and while the product is almost
valueless as a contribution to biological research, I can scarcely
suppose that it aids greatly in the advances of other branches of our
science.
But why is it that, with these exceptions, the consequences of the
admittedly general acceptance of a theory of evolution are so little
reflected in the systematic treatment of living things? Surely the
reason is that though the systematist may be convinced of the general
truth of the evolution theory at large, he is still of opinion that
species are really distinct things. For him there are still 'good'
species and 'bad' species and his experience tells him that the
distinction between the two is not simply a question of degree or a
matter of opinion.
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