[28] _Ann. N. Y. Acad. Sci._, 1878, I, p. 149.
[29] _Ibid_, 1912, pp. 523-8.
CHAPTER VII
LOCAL DIFFERENTIATION. _Continued_
OVERLAPPING FORMS
The facts of the distribution of local forms on the whole are consistent
with the view that these forms come into existence by the sporadic
appearance of varieties in a population, rather than by transformation
of the population as a whole. Of such sporadically occurring varieties
there are examples in great abundance, though by the nature of the
case it can be but rarely that we are able to produce evidence of a
previous type being actually superseded by the variety. When the two
forms are found co-existing in the same area they are usually recorded
as one species if intergrades are observed, and as two species if the
intergrades are absent. On the other hand when two forms are found
occupying separate areas, when, that is, the process of replacement is
completed in one of the areas, then forthwith each is named separately
either as species or subspecies. Successive observations carried out
through considerable periods of time would be necessary to establish
beyond question that the history proceeds in one way rather than
another. Such continuity of observation has for the most part never
been attempted. The kind of information wanted has indeed only been
lately recognized, and really critical collecting is a thing of only
the last few decades. The methods of the older collectors, who aimed at
bringing together a few typical specimens of all distinct forms, are of
little service in this class of inquiry, which is better promoted by the
indiscriminate collection of large numbers of common forms from many
localities. When this has been done on a comprehensive scale we shall be
in a position to form much more confident judgments as to the general
theory of evolution.
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