Some little work of the kind has however been done and the results are
already of great value. Seeing that the differentiation of local forms
is only made possible by isolation, it necessarily happens that the
collector finds one form in one locality and another in a distinct
locality, and there is no evidence as to the behaviour which the two
representative species might exhibit if they came into touch with each
other. In the most familiar examples of such distinction each inhabits
an island, completely occupying it to the exclusion of any other similar
form. It can only be when the two representative species occupy parts
of a continental area connected with each other by regions habitable
for the organism in question, that there is a chance of seeing the two
forms in contact. Often also, even where this condition is satisfied,
the habits, social organisation, or some other special cause may
act as a barrier which prevents the distinguishable forms from ever
coming into such complete contact as to interbreed or to behave as a
genetically continuous race. When genetic continuity is ensured by a
constant diffusion of the population over the whole area which they
inhabit there will manifestly be no formation of local races. The
practical uniformity, for example, of so many species of birds which
inhabit widely extended ranges of Western Europe is doubtless maintained
by such constant diffusion. When, as in the case of the Falcons, many
localities have peculiar forms, the fact may be taken as conclusive
evidence that there is little or no diffusion; and when we find in such
a species as the Goldfinch that in spite of migratory fluctuations there
are nevertheless geographical races fairly well differentiated, it may
similarly be inferred that these fluctuations habitually move up and
down on paths which do not intermingle. There are however a few examples
of animals, not given to much irregular wandering, which occupy a wide
and continuous range of diversified country and are differentiated as
local races in two or more districts, though the distinct races meet
in intervening areas. Of these the most notorious illustration which
has been investigated with any thoroughness is that of the species of
_Colaptes_ (Woodpeckers) known in the United States as Flickers. The
study of the variations of these forms, made by J. A. Allen[1] is an
admirable piece of work, with which every student of variation and
evolutionary problems should make himself familiar. The two forms with
which we are most concerned are known as _C. auratus_ and _C. cafer_,
and are very strikingly different in appearance. In size, proportions,
general pattern of colouration, habits, and notes, the two are alike,
but they differ in the following seven respects as stated by Allen.
_Auratus_ _Cafer_
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