cases the quills may be mostly yellow with a few red or orange
quills intermixed, or red with a similar mixture of yellow. A
bird may have the general colouration of true _cafer_ combined
with a well-developed nuchal crescent, or nearly pure _auratus_
with the red malar stripes of a _cafer_. Sometimes the body
plumage is that of _C. auratus_ with the head nearly as in pure
_cafer_, or exactly the reverse may occur. Or we may have the
general plumage as in _cafer_ with the throat and crown as in
_auratus_, and the malar stripe either red or black, or mixed
red and black, and so on in almost endless variations, it being
rare to find, even in birds of the same nest, two individuals
alike in all their features of colouration. Usually the first
trace of _cafer_ seen in _auratus_ manifests itself as a mixture
of red in the black malar stripe, either as a few red feathers,
or as a tipping of the black feathers with red, or with merely
the basal portion of the feathers red. Sometimes, however, there
is a mixture of orange or reddish quills, while the malar stripe
remains normal. In _C. cafer_ the traces of _auratus_ are usually
shown by a tendency to an incipient nuchal crescent, represented
often by merely a few red-tipped feathers on the sides of the
nape; at other times by a slight mixture of black in the red
malar stripe."
Such a state of things accords very imperfectly with expectations under
any received theory of Evolution. As in some of the instances discussed
in the first chapter we have here two fairly definite forms, nearly
allied, which on any evolutionary hypothesis must have been evolved
either the one from the other, or both from a third form at a time not
very remote from the present, as time must be measured in evolution. Yet
though intermediates exist in some quantity, no one can for a moment
suggest that they are that definite intermediate from which _auratus_
and _cafer_ descend in common. One cannot imagine that the immediate
ancestor of these birds was a mosaic, made up of asymmetrical patches
of each sort: but that is what many of the intermediates are. It is not
much easier to suppose the ancestor to have been a nondescript, with a
compromise between the developed characters of each, with quills buff,
malar stripes neither black nor red, with a trace of nuchal crescent,
and so on. Such Frankenstein-monsters have played, a considerable part
in the imaginations of evolutionary philosophers, but if it were true
that there was once a population of these monsters capable of successful
existence, surely they should now be found as a population occupying the
neutral zone between the two modern forms. Yet, though much remains to
be done in clearing up the facts, one thing is certain, namely that the
neutral zone has not a definite and normally intermediate population,
but on the contrary it is peopled by fragments of the two definite types
Public-domain text, read in full here on John Shaqi.
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