and miscellaneous mongrels between them.
On the other hand, one cannot readily suppose that either form was
the parent of the other. The process must have involved both addition
and loss of factors, for whatever hypothesis be adopted, such changes
must be supposed to have occurred. A careful statistical tabulation of
the way in which the characters are distributed in the population of
the mixed zone would be of great value, and till that has been done
there is little that can be said with certainty as to the genetics of
these characters. In the collection of Dr. Bishop of New Haven I was
very kindly allowed to examine a sample, all taken at random, near
together, in Saskatchewan. There were females 4 adult, 2 young; males
4 adult and 5 young. This number, though of course insufficient, is
enough to give some guide as to the degree of definiteness which the
characters generally show in their variations. Of the 15 birds, 8 had
simply yellow quills; 2 had red; 1 was almost red but had one yellow
tail-quill; 3 were intermediate and 1 was buff. As regards the malar
patch, which can only be determined properly in the adult males, 1 was
red, 1 was approximately red, 2 intermediate. As to nuchal crescent 4
females had none, 2 females very slight; 7 males had it, 1 had only a
slight crescent, and 1 had none. In point of quills therefore 10 were
definite out of 15; in point of crescent, 11 were definite out of 15;
and in point of malar patch 1 only was definite out of 4. The last is a
feature directly dependent on age and so counts for less, but as regards
the other two features there is some indication that the factors show
definiteness in their behaviour. It must be remembered that we have no
knowledge what the heterozygous form may be, and in the case of red
and yellow it is probably a reddish buff. The patch-works are no doubt
to be compared with other well-known pied forms, and in these we must
suppose the active factor broken up, which it probably can be very
easily. The asymmetry, which Allen notices as so marked a feature, in
the distribution of the red and yellow quills of the tail especially,
recalls that of the black markings in the pied Canaries. As is well
known to students of variations _some_ pigment-factors in _some_ animals
are apparently uncontrolled by symmetry, while in other specific cases
symmetry is the rule. On the other hand the blackness or redness of
the malar patches is, I think, as a rule nearly symmetrical. It should
be mentioned that two of Dr. Bishop's young birds belonged to the same
nest, one a female with _red_ quills, the other a male with _yellow_.
Both are without crescent.
Public-domain text, read in full here on John Shaqi.
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