As to the question whether certain combinations of characters occur
with special frequency, the evidence is insufficient to give a definite
answer. Among all the birds I have seen in America or in England I
have not yet found one having the malar patches black without any
nuchal crescent. Of Dr. Bishop's 8 adults not one, however, showed the
combination of the three chief features normal for _auratus_ or for
_cafer_.
Besides the two forms that we have hitherto considered, several other
local types exist, and these throw some further light on the problem.
Of these the most important in this connexion is _chrysoides_, which
inhabits the whole of southern California and the mainland opposite.
This remarkable form is as Allen says, very different from _auratus_
except that it has the quills yellow like _auratus_, not red like
_cafer_. So that we find here in the extreme west of the whole
distribution a type agreeing in one of its chief features with the
eastern type. Between this and _cafer_ intergrades have, according to
Allen, not been found. The relations of this _chrysoides_ are, Allen
thinks, rather with _mexicanoides_, a southern, smaller race with
colours more intense, which inhabits Guatemala, but however that may
be, it must be regarded as a _cafer_ which has lost its red quills. The
island of Guadeloupe off Lower California has an island form. Beyond the
other side of the continent there is also an island form of _auratus_,
inhabiting Cuba, so that clearly the yellow quills can extend into the
tropics.
The above account is in many respects incomplete, but it suffices to
give an outline of the chief facts. The whole problem is complicated by
the undoubted effects of an uncertain amount of migration, and in many,
perhaps all, districts, the winter population differs from the summer
population of the same localities. The existence of these seasonal ebbs
and flows is now well known to ornithologists, and most of the bird
species of temperate regions are subject to them.
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