Before speaking of the distribution in other parts of France and in
Europe generally I will briefly state the results of my breeding
experiments. The work was done many years ago before we had the
Mendelian clue, and it is greatly to be hoped that some one will
find opportunities of repeating it. Crossing the English and the
thoroughly southern type the families produced agree entirely with
the intermediates of Brittany and the Loire. Reciprocals are alike.
Of F_{2} I only succeeded in raising very few and of those that I had
(about 30) nearly all were intermediate in character, though perhaps
rather less uniform than F_{1}. One family alone, containing only 4
specimens, had one _egerides_, and three fulvous intermediates. As the
case stands alone I hesitate whether or not to suppose it due to some
mistake. Moreover from F_{1} crossed back with the respective parental
types I had fairly long series, especially from F_{1} × the southern
type, and looking at these families I cannot see any clear evidence
of segregation. On the contrary, I think that though there are slight
irregularities, they would, taken as a whole, be classed as coming
between the intermediate type and the extreme form used as the second
parent. This at least is true when the second parent was of the southern
type.
On this evidence I have regarded the case as one in which there is
no good evidence of segregation and as conforming most nearly with
the conventional view of gradual transition in response to climatic
influences. Such influence must however be indirect; for I reared five
generations of the northern type in England, and these, though they
included several abnormal-looking specimens in the last generation and
then died out, did not show any noticeable change from the fulvous
colour of the wild type. Merrifield[8] also found that heat applied to
pupae of the northern type produced no approach to the southern type.
Public-domain text, read in full here on John Shaqi.
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