Is it not time to abandon these fanciful expectations which are never
realised? Everywhere both among animals and plants does the problem of
specific difference reiterate itself in the same form. In view of such
facts as I have related and might indefinitely multiply, the fixity of
specific characters cannot readily be held to be a measure of their
economic importance to their possessors. The incidence of specific
fixity is arbitrary and capricious, sometimes lighting on a feature or
a property which can be supposed to matter much, but as often is it
attached to the most trifling of superficial peculiarities.
The incidence of _variability_ is no less paradoxical, and without
investigation of the particular case no one can say what will be found
to show much or little variability. The very characteristic which in
one species may exhibit extreme variability may in an allied species
show extreme constancy. Illustrations will occur to any naturalist, but
nowhere is this truth more strikingly presented than in the British
Noctuid Moths. Many are so variable that, in the common phrase,
"scarcely two can be found alike," while others show comparatively
slight variation. It need scarcely be remarked that, in the instances
I have in mind, the evidence of great variability is in no way due to
the abundance with which the particular species occurs, for common
species may show constancy, and less abundant species may show great
variability. The polymorphism seems to be now at least a general
property of the variable species, as the fixity is a property of the
fixed species. In illustration I may refer to the following examples.
_Dianthoecia capsincola_ is a common and widely distributed moth which
feeds on _Lychnis_. It shows little variation. _Dianthoecia carpophaga_
is another species which feeds chiefly on _Silene_. Its habits are
very similar to those of _capsincola_. Like that species it has a wide
geographical range and is abundant in its localities, but in contrast
to the fixity of _capsincola_, _carpophaga_ exhibits a complex series
of varieties. _Agrotis suffusa_ (= _ypsilon_) is a moth widely spread
through the southern half of England. It is very constant in colour and
markings. _Agrotis segetum_ and _tritici_ are excessively variable both
in ground colour and markings, being found in an immense profusion of
dissimilar forms throughout their distribution. Of these and several
other species of _Agrotis_ there are many named varieties, some of
which have by various writers been regarded as specifically distinct.
Of the genus _Noctua_ many species (e. g. _festiva_) show a similar
polymorphism, but _N. triangulum_, though showing some variation in
certain respects, is usually very constant to its type, and the same is
true of _N. umbrosa_.
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