This fact is difficult to reconcile with the conventional view that
specific type is directly determined by Natural Selection and that
the precision with which a species conforms to its pattern is an
indication of the closeness of that control. Anyone familiar with the
characteristics of Moths will agree that the Noctuids, Geometrids and
Tortricids are creatures whose existence depends in some degree on the
success with which they can escape detection by their enemies in the
imaginal state. We are therefore not surprised to find that some species
of these orders exhibit definite geographical variation in conformity
with the character of the ground, which may reasonably be supposed to
aid in their protection. If this were all, there would be nothing to
cause surprise. We might even be disposed to allow that variability
might contribute to the perpetuation of animals so situated, on the
principle that among a variety of surroundings some would probably be
in harmony with the objects on which they rest. But we cannot admit
the plausibility of an argument which demands on the one hand that
the extreme precision with which species A adheres in the minutest
details of its colour and pattern to a certain type shall be ascribed
to the protective fitness of those details, and on the other hand that
the abundant variability of species B shall be ascribed to the same
determination. If it is absolutely necessary for A to conform to one
type how comes it that B may range through some twenty distinct forms,
any two of which differ more from each other than the regular species
of many other genera? The only reply I can conceive is a suggestion
that there _may_ be some circumstance which differentiates the various
classes of cases, that the exigencies of the fixed species _may_ be
different from those of the variable. Those who make such appeals to
ignorance do not always perhaps realise whither this course of reasoning
may lead. If admissible here the same argument would lead us to suggest
that because albino moles have for an indefinite period occurred on a
certain land near Bath there may be something in the soil or in the
conditions of life near Bath which requires a proportion of albinos in
its mole population. Or again, because the butterfly _Thais rumina_
in one locality, Digne in the south of France, has a percentage of
individuals of the variety _Honoratii_ (with certain normally yellow
spots on the hind wing coloured bright red) and nowhere else throughout
its distribution, that therefore we may suggest that there is some
difference in the condition of life at Digne which makes the continuance
of _Honoratii_ there possible and beneficial.
Public-domain text, read in full here on John Shaqi.
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