Our tissues therefore are like a beach composed of sands of
different kinds, and different kinds of sands may show distinct and
interpenetrating ripples. When the essential analogy between these
various classes of phenomena is perceived, no one will be astonished at,
or reluctant to admit, the reality of discontinuity in Variation, and if
we are as far as ever from knowing the actual causation of pattern we
ought not to feel surprised that it may arise suddenly or be suddenly
modified in descent. Biologists have felt it easier to conceive the
evolution of a striped animal like a zebra from a self-coloured type
like a horse (or of the self-coloured from the striped) as a process
involving many intergradational steps; but so far as the _pattern_ is
concerned, the change may have been decided by a single event, just as
the multitudinous and ordered rippling of a beach may be created or
obliterated at one tide.
[Illustration: FIG. 1. Tusk of Indian elephant, showing an abnormal
segmentation.]
This point is well illustrated by the tusk of an Indian elephant which
I lately found in a London sale-room. This tusk is by some unknown
cause, presumably a chronic inflammation, thrown up into thirteen
well-marked ridges which closely simulate a series of segments (Fig. 1).
Whatever the cause the condition shows how easily a normally unsegmented
structure may be converted into a series of repeated parts.
The spread of segmentation through tissues normally unsegmented is very
clearly exemplified in the skates' jaws shown in Fig. 2. The right
side of the upper figure shows the normal arrangement in the species
_Rhinoptera jussieui_, but the structure on the left side is very
different. The probable relations of the several rows of teeth to the
normal rows is indicated by the lettering, but it is evident that by
the appearance of new planes of division constituting separate centers
of growth, the series has been recast. The pattern of the left side is
so definite that had the variation affected the right side also, no
systematist would have hesitated to give the specimen a new specific
name. The other two drawings show similar variations of a less extensive
kind, the nature of which is explained by the lettering of the rows of
teeth.
[Illustration: FIG. 2. Jaws of Skates (_Rhinoptera_) showing meristic
variation. (For a detailed discussion see _Materials for the Study of
Variation_, p. 259.)]
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