It is a feature common to these and to all other rhythmical repetitions
produced by purely mechanical forces that there is resemblance between
the members of the series, and that this similarity of conformation may
be maintained in most complex detail. When however in the mechanical
series some of the members differ from the rest we have no difficulty
in recognising that these differences--which correspond with the
differentiations of the organic series--are due to special heterogeneity
in the conditions or in the materials, and it never occurs to us to
suppose that all the members must have been primordially alike. For
example, in the case of ripple-marks on the sand, which I choose as one
of the most familiar and obvious illustrations of a repeated series
due to mechanical agencies, if we notice one ripple different in form
from those adjacent to it, we do not suppose that this variation must
have been brought about by deformation of a ripple which was at first
formed like the others, but we ascribe it to a difference in the sand at
that point, or to a difference in the way in which the wind or the tide
dealt with it. We may press the analogy further by observing that in as
much as such a series of waves has a beginning and an end, it possesses
polarity like that of the various linear series of parts in organisms,
and even the formation of each member must influence the shape of its
successor. Since in an organism the beginning and end of the series
are always included, some differentiation among the repetitions must
be inevitable. If therefore it be conceded, as I think it must, that
segmentation and pattern are the consequence of a periodic process we
realize that it is at least as easy to imagine the formation of such a
series of parts having family likeness combined with differentiation
as it would be to conceive of their arising primordially as a series
of identical repetitions. The suggestion that the likenesses which we
now perceive are the remains of a still more complete resemblance is a
substitution of a more complex conception for a simpler one.
The other question raised by the problem of Serial Homology is how far
there is a correspondence between individual members of series when
the series differ from each other either in the number of parts, or in
the mode of distribution of differentiation among them. Students, for
example, of vertebrate morphology debate whether the _n_th vertebra
which carries the pelvic girdle in Lizard A is individually homologous
with the _n_ + _x_th vertebra which fulfils this function in Lizard B,
or whether it is not more truly homologous with the vertebra standing in
the _n_th ordinal position, though that vertebra in Lizard B is free.
Public-domain text, read in full here on John Shaqi.
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