It may be urged that no feature as yet enables us to perceive wherein
lies the primary distinction which determines such variation, whether it
is due to a difference in the dividing forces or in the material to be
divided. If for instance we were to imitate such a series of segments
by pressing hanging drops of a viscous fluid out of a paint-tube by
successive squeezes, the number of times the tube is contracted before
it is empty will give the number of the segments, but their size may
depend either on the force of the contractions or on the capacity of
the tube, or on various other factors. Nevertheless in the case of the
variation of terminal members, whatever be the nature of the rhythmical
impulse which produces the series of organs, the elevation of the
normally terminal member in correspondence with the addition of another
is what we should expect.
If the organism acquired its full size first and the delimitation of
the parts took place afterwards, there might be some hope that the
resemblance between living patterns and those mechanically caused by
wave-motion might be shown to be a consequence of some real similarity
of causation, but in view of the part played by growth, appeal to these
mechanical phenomena cannot be declared to have more than illustrative
value. Similarly in as much as living patterns appear, and almost
certainly do in reality come into existence by a rhythmical process,
comparisons of these patterns with those developed in crystalline
structures, and in the various fields of force are, as it seems to me,
inadmissible, or at least inappropriate.
However their intermittence be determined, the rhythms of division must
be looked upon as the immediate source of those geometrically ordered
repetitions universally characteristic of organic life. In the same
category we may thus group the segmentation of the Vertebrates and of
the Arthropods, the concentric growth of the Lamellibranch shells or of
Fishes' scales, the ripples on the horns of a goat, or the skeletons of
the Foraminifera or of the Heliozoa. In the case of plant-structures
Church[4] has admirably shown, with an abundance of detail, how on
analysis the definiteness of phyllotaxis is an expression of such
rhythm in the division of the apical tissues, and how the spirals
and "orthostichies" displayed in the grown plant are its ultimate
consequences. The problem thus narrows itself down to the question of
the mode whereby these rhythms are determined.
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