[Illustration: FIG. 11. Diagrams of the geometrical relations which are
generally exhibited by extra pairs of appendages in Arthropoda. The
sections are supposed to be those of the apex of a tibia in a beetle.
_A_, anterior, _P_, posterior, _D_, dorsal, _V_, ventral. _M_^{1},
_M_^{2} are the imaginary planes of reflexion. The shaded figure is in
each case a limb formed like that of the other side of the body, and the
outer unshaded figures are shaped like the normal for the side on which
the appendages are. On the several radii are shown the extra pairs in
their several possible relations to the normal from which they arise.
The normal is drawn in thick lines in the center.]
Next the geometrical relations of the halves of the supernumerary pair
are determined by the position in which they stand in regard to the
original appendage. These relations are best explained by the diagram
(Fig. 11), from which it will be seen that the two supernumerary
appendages stand as images of each other; and, of them, that which is
adjacent to the normal appendage forms an image of it. Thus if the
supernumerary pair arise from a point on the dorsal surface of the
normal appendage, the two _ventral_ surfaces of the extra pair will
face each other. If they arise on the anterior surface of the normal
appendage, their morphologically posterior surfaces will be adjacent,
and so on.
These facts give us a view of the relations of the two halves of a
dividing bud very different from that which is to be derived from the
exclusive study of normal structures. Ordinary morphological conceptions
no longer apply. The distribution of the parts shows that the bud or
rudiment which becomes the supernumerary pair may break or open out in
various ways according to its relations to the normal limb. Its planes
of division are decided by its geometrical relations to the normal body.
Especially curious are some of the cases in which the extra pair are
imperfectly formed. The appearance produced is then that of two limbs
in various stages of coalescence, though in reality of course they are
stages of imperfect separation. The plane of "coalescence" may fall
anywhere, and the two appendages may thus be compounded with each other
much as an object partially immersed in mercury "compounds" with its
optical image reflected from the surface.
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