the result shows the influence of the major component on the small
piece, causing it to produce a head and not a tail at its aboral end,
but I think that this conclusion would be erroneous, because it seems
much more probable that we have here a case of heteromorphosis, similar
to that in _Planaria lugubris_, and that the result depends entirely on
the action of the smaller component. It is hardly possible to
demonstrate that this is the correct interpretation, since if a small
piece of this size is isolated it dies before it regenerates. The result
is paralleled, however, by the regeneration of a tail at the anterior
surface of a posterior piece.
The process of grafting has long been practised with plants, but the
experiments were made more for practical purposes than to study the
theoretical problems involved. Vöchting has, however, carried out a
large number of well-planned experiments. He finds that a stem can be
grafted upon a root, and a root upon a stem, a leaf upon a stem or upon
a root. Even an entire plant can be grafted upon another. The results
show, however, in general, that, whatever the new position may be, the
graft retains its morphological characters--a shoot remains a shoot, a
root is always a root, and a leaf a leaf. Vöchting concludes that there
is in the plant no principle or organization that conditions an
unchangeable arrangement of the main organs. “The inherited order of the
parts, acquired apparently on physiological grounds, may be altered by
the experimentator; it is possible for him to change the position of the
building blocks within a wide range without endangering the life of the
whole.” “It is essential, however, for the success of the experiment
that the grafted parts, or tissues, retain their normal orientation. If
this condition is not fulfilled there may take place, it is true, a
union of the parts, but sooner or later disturbances set in.” Vöchting
transplanted pieces in abnormal positions, sometimes reversing the long
axis of the grafted piece, sometimes the radial axes, and sometimes both
together. In some cases this led to the formation of swellings that
interfered with the nourishment but carried with it no further
consequences. In other cases the changes went so far that the vital
processes were interfered with. At times an incomplete union took place
between the parts; at others, even though the first union was perfect,
death later ensued.
Public-domain text, read in full here on John Shaqi.
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