On the other hand, when similar pieces were grafted with their original
orientation, a perfect union took place and the piece became a part of
the stock. The results establish, Vöchting claims, that every part and
every portion of a part has a polar orientation in one direction, and
furthermore, in a body having a radially symmetrical form, there is also
a radial polarization; that is, the inner side of each part is different
from the outer side of the same surface, even though no such difference
is apparent to us. The properties of the tissue-complex rest, in the
last analysis, on that of the cells; the properties of the whole being
only the sum total of the properties of its elements, so that we may say
that every living cell of the root is polarized, not only
longitudinally, but also radially; each has a different apical and root
pole, a different anterior and posterior pole, and also right and left
polar relations. These results, deduced from the experiments in
grafting, lead Vöchting to formulate the following rule: “Like poles
repel, unlike poles attract.” This rule is the same as the law of the
magnet. In fact, Vöchting states that the root and the stem relations
show a remarkable resemblance, despite many differences, to a magnet. If
the magnet is broken into pieces it may be reunited by bringing unlike
poles together, but not by uniting like poles; the same statement holds
for the root and the stem.
Exception may be taken, I believe, to parts of Vöchting’s conclusions,
especially in the light of the recent experiments in grafting in
animals. It is by no means to be granted without further demonstration
that the properties of the whole organism are only the sum-total of the
action of the individual cells. If, as seems to be the case, the cells
are organically united into a whole, the properties of this whole may be
very different from the sum of the properties of the individual cells,
just as the properties of sugar are entirely different from the sum of
the properties of carbon, hydrogen, and oxygen.
Public-domain text, read in full here on John Shaqi.
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