The Trilobites, after appearing in a great variety of generic and
specific forms, and playing a most important part in their time,
were not destined to continue beyond the Carboniferous period, and
before that time they were beginning to give place to the Limuli,
King-crabs, or Horseshoe-crabs, a few species of which continue on our
coasts until the present time. In this limited duration the Trilobites
present a strange contrast to certain shrimp-like Crustaceans, their
contemporaries (the Phyllopods), which very closely resemble some
still extant, and the same remark applies to swarms of little bivalve
Crustaceans (Ostracods), which are still represented by hosts of modern
species both in the sea and in the fresh waters. There is, however,
a remarkable group of shrimp-like Crustaceans, represented in the
modern world by only a few small species, which in the Cambrian age
attained greater size, and constitute a very generalized type combining
characters now found in lower and higher groups of Crustacea.
_Hymenocaris vermicauda_ of Salter (Fig. 3) may serve to illustrate one
of these primitive forms.
[Illustration: Fig. 3.--_Hymenocaris vermicauda_, Salter.
A Lower Cambrian Shrimp of generalized type. (After Salter.)]
In point of fact, as Dr. Henry Woodward has shown in an able
presidential address delivered to the Geological Society in 1895, at
the base of the Lower Cambrian we still have several distinct groups
of Crustacea; and if with some we were to hold them as traceable to
one original form or to a worm-like ancestor, we must seek for this
far back in those pre-Cambrian rocks in which we find no Crustaceans
whatever. There is, it is true, no good reason to demand this; for
whatever the cause, secondary or final, which produced any form of
Crustacean in the Lower Cambrian, it might just as well have produced
several distinct forms. Evolutionists seem to be somewhat unreasonable
in demands of this kind, for any cause capable of originating a new
form of living being, might have been operative at the same time in
different localities and under somewhat diverse conditions, and may
also have acted at different times. All imaginary lines of descent of
animals are more or less subject to this contingency; and this may
partly account for the great diversity in the lines of affiliation
presented to us by evolutionists, which may in part have a basis in
fact in so far as distinct varietal and racial forms are concerned, but
may just as likely be entirely fallacious in the case of true species.
In any case, in the lowest rocks into which we can trace Crustacea, we
have already probably five of the orders into which their successors
in the modern seas are divided by zoologists; and this is certainly a
singular and suggestive fact, the significance of which we shall be
better prepared to understand at a later stage of our investigation.
Public-domain text, read in full here on John Shaqi.
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