We miss here the two great Molluscan groups of the creeping Sea-snails
like the limpet and whelk, and of the ordinary bivalves like the
oyster and cockle. Both are present in the lowest Cambrian, though in
small numbers compared with their present abundance. Possibly they had
not yet appeared in the Etcheminian Sea, though the muddy and sandy
bottoms, evidenced by its slates and sandstones, would seem to have
afforded favourable habitats, and warrant the expectation that species
may yet be found.
The case was different with the little group of the Lamp-shells, or
Brachiopods. These creatures, somewhat resembling the ordinary bivalves
in their shelly coverings, were very dissimilar in their internal
structure, and once settled on the bottom they were attached for life,
not having even the limited means of locomotion possessed by the
Sea-snails and common bivalves. They collected their food wholly by
means of currents of water produced by cilia, or movable threads, on
arms or processes within their shells. In this they resembled the young
or embryo stages of some of the more ordinary Mollusks, though they are
so remote from these in their adult condition that they have usually
been placed in a distinct class, and some naturalists have thought
it best to separate them from the Mollusks altogether. Their history
is peculiar. Coming into existence at a very early date, they became
very abundant in early Palæozoic times, then gradually gave place to
the ordinary bivalves, and in the modern seas are represented by very
few species. Yet while in the middle period of their history they are
represented by very many peculiar specific and generic forms. Some of
the earliest types, like Obolus and Lingula, persist very long, and
the latter has continued without change from the Early Cambrian to the
Modern period.
The great group of the Sea-stars and Sea-urchins appears only in a
few of its lower forms, and seems to be the only class represented by
embryonic types. The coral animals are absent, so far as known. The
Jelly-fishes and their allies cannot be preserved as fossils, but some
peculiar markings, at one time regarded as plants, are now supposed to
be trails made by the tentacles of creatures of this kind moving over
muddy bottoms. A few spicules indicate Sponges, and the ubiquitous
groups of the marine Protozoa, the Foraminifera and the Radiolaunus,
are represented by shells scarcely distinguishable from those of modern
species. The great and peculiar forms represented at this early time by
Cryptozoon and its allies seem long ago to have perished, and we shall
have to return to them in a later stage of our inquiry.
Public-domain text, read in full here on John Shaqi.
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