Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
103. _The Physical Properties of the Skeleton._--The skeletons of all
Radiolaria are characterised pre-eminently by a high degree of _firmness_,
which fits them to serve as protective and supporting apparatus. This is
obvious in the case of the pure siliceous shells of the Polycystina; but
the acanthin framework of the ACANTHARIA also possesses a degree of
stiffness but little inferior, whilst the silicate skeletons of the
PHÆODARIA seem on the whole to be not so firm. The hollow skeletal tubes of
the last-named, which are filled with gelatinous material, are very brittle
on account of the delicacy of their walls. Their _elasticity_ also is very
small, whilst that of the acanthin spines is considerable. The thin long
needles of many ACANTHARIA are very elastic, as are also the bristle-like
siliceous spicules of many SPUMELLARIA. The _refractive power_ of the
skeleton in the various legions is very different, depending upon the
chemical constitution. The siliceous skeleton of the Polycystina
(SPUMELLARIA and NASSELLARIA) and the silicate skeleton of the PHÆODARIA
have the same refractive index as glycerine, and hence become invisible
when mounted in that fluid; they then become visible only on addition of
{lxxi}water, and are clearer in proportion to the quantity of water which
is added. The refractive index of acanthin is, however, very different from
that of glycerine, so that the skeletons of ACANTHARIA are readily visible
when mounted in this fluid. In water, the skeletons of all Radiolaria
appear about equally refractive, as also in Canada balsam. The substance of
the skeleton appears almost entirely hyaline, colourless, and transparent.
Very rarely it is faintly coloured (in some ACANTHARIA). A cloudy opaque
constitution is seen in some PHÆODARIA (especially in the "porcellanous
shells" of Tuscarorida and Circoporida, Pls. 100, 114-117); when dried,
these appear by reflected light milky-white or yellowish-white; the cause
of this opacity lies partly in the peculiar "cement-like structure" of
these porcellanous shells, partly in their fine porosity, and the minute
air-bubbles contained in their thick walls.
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