Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
104. _The Elementary Structure of the Skeleton._--The general constitution
of the skeleton--or more accurately expressed, of the morphological
elements of which the skeleton consists--is of such a nature that it may be
termed structureless. Both the organic acanthin skeletons of the ACANTHARIA
and the silicate skeletons of the PHÆODARIA, as well as the inorganic
siliceous skeletons of the SPUMELLARIA and NASSELLARIA, appear under the
microscope perfectly homogeneous, transparent, colourless, and crystalline.
Only very rarely do they show traces of a concentric striation, which
arises from the deposition of the skeletal substance in layers; as, for
example, the thick spines of some PHÆODARIA (Pls. 105-107, &c.). Some of
the PHÆODARIA, however, form an exception to this rule, inasmuch as their
partially tubular skeletal elements possess a remarkable porcellanous
structure. In the tubular or Cannoid skeleton, which occurs in most
CANNOPYLEA, the lumen of the thin-walled flinty tube is filled with jelly,
and frequently a thin siliceous thread runs in its axis, and is connected
with the wall by transverse threads (§§ 127, 139). The elementary structure
of the opaque porcellanous shells, which distinguish the two families
Circoporida (Pls. 114-117) and Tuscarorida (Pl. 100), is quite peculiar.
Numerous fine siliceous spicules lie scattered irregularly in a finely
granular or porous matrix.
105. _Complete and Incomplete Lattice-Shells._--In the great majority of
Radiolaria (in all four legions) the skeleton has the form of a delicate
lattice-shell or a receptacle in which the central capsule is enclosed. In
a small minority, however, this is not the case. The skeleton then consists
only of isolated rigid pieces (radial or tangential spicules), or of a
simple ring (sagittal ring of the #Stephoidea#), or of a basal tripod with
or without a loose tissue of trabeculæ, &c. (#Plectoidea#); the central
capsule is then not surrounded by a special latticed receptacle, but only
rests upon the skeletal trabeculæ. According to these different
arrangements, two principal groups or sublegions may be distinguished in
each legion, of which one set (Cataphracta) are characterised by a complete
{lxxii}lattice-shell, whilst the others (Aphracta) are without it. The
RADIOLARIA APHRACTA, then, or Radiolaria without a complete skeleton, are
the #Collodaria# (p. 9), the #Acanthometra# (p. 725), the #Plectellaria#
(p. 895), and the #Phæocystina# (p. 1543). On the other hand, the
RADIOLARIA CATAPHRACTA, or Radiolaria with a complete skeleton, are the
#Sphærellaria# (p. 49), the #Acanthophracta# (p. 791), the #Cyrtellaria#
(p. 1015), and the #Phæocoscina# (p. 1590).
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