Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
112. _The Skeleton of the Phæodaria._--The skeleton of the PHÆODARIA or
CANNOPYLEA is always extracapsular, usually consists of a silicate of
carbon (more rarely of pure silica), and in the majority of the legion is
composed of hollow cylindrical tubes, whose siliceous wall is very thin,
and whose lumen is filled with gelatinous material (§ 127). The manifold
and remarkable skeletal forms occurring in this legion are not
monophyletic, since they cannot be derived from a common stem-form; they
are, on the contrary, polyphyletic, various skeletonless PHÆODARIA
(Phæodinida) have independently acquired skeletons of different form and
composition. The legion PHÆODARIA can be subdivided into four orders, the
skeletons of which present the following important distinctions:--(1) The
#Phæocystina# possess only incomplete Beloid skeletons (§ 115), composed of
many separate pieces, sometimes tangentially (Cannorrhaphida, Pl. 101),
sometimes radially arranged (Aulacanthida, Pls. 102-105). (2) The
#Phæosphæria# form Sphæroid skeletons (§ 116), usually only a simple
lattice-shell without special aperture (Pls. 106-111); two concentric
shells united by radial bars occur only in the Cannosphærida (Pl. 112). (3)
The #Phæogromia# are distinguished by the formation of a simple Cyrtoid
skeleton (§ 123) resembling that of the Monocyrtida; the monothalamus
lattice-shell is usually ovoid or helmet-shaped, more rarely polyhedral or
almost spherical; a vertical main axis can always be distinguished, at the
basal pole of which is an aperture usually armed with teeth or spines (Pls.
99, 100, 113-120). (4) The #Phæoconchia# are distinguished from all other
Radiolaria by the possession of a bivalved shell like that of the
Conchifera; the two valves of this Conchoid skeleton must be distinguished
as dorsal and ventral, as in the Brachiopoda (Pls. 121-128). The fifteen
families of PHÆODARIA which are arranged in the four orders just mentioned,
present such great differences among themselves, that the skeleton must be
regarded as probably polyphyletic even within the limits of each order.
113. _Types of Skeletal Formation._--No less than twelve different
principal forms may be distinguished as morphological types of the
formation of the skeleton in the Radiolaria; some of these are peculiar to
a single legion or even to a smaller group; but sometimes the same form
occurs in several legions. Some types occur only in an isolated manner,
independently of the others, but most exist in various combinations with
other types. Of the twelve described below the Conchoid and Cannoid occur
only in the PHÆODARIA; the Plectoid and Circoid only in the NASSELLARIA;
the Astroid only in the ACANTHARIA; the remaining seven types are found in
several legions in the same form and hence are polyphyletic.
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