Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
{lxxvii}114. _The Astroid Skeleton._--Under the name "Astroid" we place the
peculiar star-shaped skeletons of the ACANTHARIA in opposition to those of
all other Radiolaria, for they are separated from them not only
fundamentally by reason of the chemical nature of their substance
(Acanthin, § 102), but also by their centrogenous origin, and the resulting
stellate form (Pls. 129-140). The ACANTHARIA are the only Radiolaria in
which the skeleton arises within the central capsule by the formation of
numerous rays or radial spines of acanthin which project on all sides from
the centre. Originally these are united at this point, their conical or
pyramidal points meeting and being supported one upon another. In the great
majority of ACANTHARIA this loose apposition is constant, so that when the
soft parts are destroyed the skeleton falls to pieces. Only in a few forms
in this legion are the central ends of the spines fused so that the whole
skeleton forms a connected star (_Astrolithium_). The small group
Chiastolida (or Acanthochiasmida) is characterised by the fact that the two
rays which are opposite to one another in each axis unite and form a
diametral bar. The skeleton is almost always composed of twenty radial
spines, which are regularly disposed (Icosacantha), only in the small
primitive group #Actinelida# is the number variable (Adelacantha, § 110).
115. _The Beloid Skeleton._--As Beloid or spicular skeletons are grouped
together all those which consist of several disconnected portions; these
always lie outside the central capsule, either within the calymma or on its
surface. Such extracapsular Beloid skeletons are entirely wanting in the
ACANTHARIA and NASSELLARIA; they occur only in the #Beloidea# among the
SPUMELLARIA, and in the #Phæocystina# among the PHÆODARIA; the individual
Beloid portions of the former are solid, those of the latter hollow. In
both groups the simplest forms of the separate portions are simple
unbranched needles (_Thalassosphæra_, _Thalassoplancta_, _Physematium_,
_Belonozoum_, among the SPUMELLARIA; _Cannobelos_ and _Cannorrhaphis_ among
the PHÆODARIA); usually these spicules are disposed tangentially over the
surface of the calymma. Among the #Beloidea# branched spicules occur more
commonly than these simple ones; they are either stellate (with many rays
united in a centre) or twin-like, with a tangential bar, from each pole of
which two or three (seldom more) radial branches project (Pls. 2, 4). Among
the PHÆODARIA the subfamily Dictyochida is characterised by the annular
shape of its Beloid portions, either simple rings, or hat-shaped or
pyramidal bodies with a latticed cap over the ring (Pl. 101, figs. 3-14;
Pl. 114, figs. 7-13). The family Aulacanthida among the PHÆODARIA, alone
possesses hollow _radial tubes_, which penetrate the whole calymma, and
project distally over its surface, whilst their proximal ends rest upon the
surface of the central capsule. Although in these cases the enclosed
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