Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
146. _The Ontogeny of the Spumellaria._--In the simplest case the
individual development in the SPUMELLARIA ceases with the _Actissa_-stage.
In all other genera of this legion diverging forms proceed from this, of
which the different growth of the three dimensive {xcvi}axes on the one
hand (§§ 44, 45), and the differentiation of the various parts of the
unicellular organism with the formation of the skeleton on the other, are
of pre-eminent significance. Even in the varying growth of the central
capsule in the different dimensions of space in the skeletonless
#Colloidea#, four different modes may be distinguished, which further, in
the corresponding development of the skeleton, furnish the basis for the
origin of the four orders of #Sphærellaria#. The most primitive and
simplest form of growth, equal extension in all directions, is found in the
spherical central capsule and the concentric spherical skeletons
(_Procyttarium_, #Sphæroidea#). When the growth of the central capsule
proceeds more rapidly in the direction of the vertical main axis than in
any other direction, the ellipsoidal or cylindrical central capsule
(_Actiprunum_) arises, and the vertically elongated skeleton of the
#Prunoidea#, which is derived from it. When, on the contrary, the growth of
the central capsule and lattice-shell is less in the direction of the
vertical main axis than in any other direction, the lenticular or discoid
central capsule (_Actidiscus_) arises, and the corresponding lenticular
shell of the #Discoidea#. Finally, even quite early in many SPUMELLARIA,
the growth of the central capsule and of the corresponding lattice-shell in
the three dimensive axes is different, and hence arise the lentelliptical
forms whose geometrical type is the triaxial ellipsoid or the rhombic
octahedron (_Actilarcus_, #Larcoidea#). Thus the origin of the four orders
of #Sphærellaria# is simply explained by a varying growth in the different
dimensive axes. The _primary_ (innermost) lattice-shell is in this legion
always _simultaneously_ developed (suddenly excreted at the moment of
lorication from the sarcodictyum). The _secondary_ lattice-shells, on the
other hand, which surround the former concentrically, and are united with
it by radial bars, arise _successively_ from within outwards.
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