Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
147. _The Ontogeny of the Acantharia._--The individual development of the
ACANTHARIA in the simplest case (_Actinelius_) stops at a point which
differs from the _Actissa_-stage only in the change of radial axial threads
into acanthin spines. In the small group #Actinelida#, their number remains
variable and usually indeterminate (Adelacantha), whilst in the great
majority of the legion (#Acanthonida# and #Acanthophracta#) the number is
constantly twenty, and those spines are regularly arranged according to the
Müllerian law in five parallel circles, each containing four crossed spines
(Icosacantha). The simplest form among these latter is _Acanthometron_,
which may be regarded both ontogenetically and phylogenetically as the
common starting-point of all the Icosacantha. Within this extensive group
variations in the length of the dimensive axes appear, similar to those
observed in the SPUMELLARIA. In the Astrolonchida and #Sphærophracta# the
central capsule remains spherical, extending equally in all directions; and
correspondingly the lattice-shell, which is excreted on the surface of the
spherical calymma, remains spherical. In the Belonaspida (just as in the
#Prunoidea#) {xcvii}this form passes over into an ellipsoid by prolongation
of one axis; on the contrary, in the Hexalaspida (as in the #Discoidea#)
the discoidal or lenticular form arises by shortening of an axis. Finally,
in the Diploconida, and in some Hexalaspida in which the growth is
different in all three dimensive axes (as in the #Larcoidea#), both the
central capsule and the shell assume the lentelliptical form. The
lattice-shell of the #Acanthophracta# is usually successive in its
development, since from each of the twenty radial spines two or four
tangential apophyses proceed, whose branches subsequently unite and combine
to form the lattice-shell. Only in the peculiar Sphærocapsida can the
pavement-like shell arise simultaneously or in a moment of lorication.
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