Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
The trizonal _Larnacilla_-shell commences by the formation of a
transverse girdle, by the union of two lateral latticed processes, which
spring right and left in the equatorial plane from the poles of the
frontal axis of a lentelliptical medullary shell (_Monozonium_, p. 633,
Pl. 9, fig. 1). This is followed by a second lateral girdle, which lies
in the frontal plane and proceeds from its lateral poles (_Dizonium_, p.
634, Pl. 9, figs. 2, 3). Finally the sagittal girdle is formed, lying in
the sagittal plane and arising from the lateral girdle on the two poles
of the main axis (_Trizonium_, p. 637, Pl. 9, fig. 4). Whilst the gaps
between the three zones of this trizonal shell remain open in the
Pylonida, in _Larnacilla_, the important primitive form of the Larnacida,
they are closed by lattice-work (Pl. 50, figs. 3-8). From this trizonal
_Larnacilla_-shell the great majority of Larcoid shells may be derived.
Such a system of zones may be repeated (Diplozonaria) or even developed a
third time (Triplozonaria, p. 632). In most #Larcoidea# the zones are
secondarily connected by lattice-work. In the Tholonida (Pl. 10) each of
the two opposite latticed wings of a zone becomes a closed dome. In the
Zonarida (Pl. 50, figs. 9-12) these domes are partially or wholly
bisected by constrictions or latticed septa which are developed in the
three dimensive planes. The Lithelida (Pl. 49, figs. 1-7) are
characterised by the fact that one of each pair of opposite latticed
processes (or half zones) grows more strongly than the other, and that
the larger completely embraces the smaller so as to form a complicated
spiral. Whilst in this case the spiral lies in a plane, in the
Streblonida (Pl. 49, figs. 8, 9) it becomes turbinoid like a gastropod
shell and forms an ascending spiral. Finally, two small families of
#Larcoidea# are characterised by quite irregular growth (a very rare
occurrence among the Radiolaria); these are the simple-chambered
Phorticida (Pl. 49, figs. 10, 11) and the many chambered Soreumida (Pl.
49, figs. 12, 13). The phylogenetic relationship of these families of
#Larcoidea# is probably very complicated and demands closer investigation
(compare pp. 599-604).
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