Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
37. _The Amphipleural Ground-Forms._--By the term amphipleural ground-forms
are to be understood those usually defined as "bilaterally radial"; their
geometrical type is a half amphithect pyramid. The best known examples of
this form in the animal kingdom are the bilateral five-rayed Echinoderms
(_Spatangus_, _Clypeaster_), in the vegetable kingdom the symmetrical
five-rayed flowers (_Viola_, _Trifolium_). The three dimensive axes have
the same relation as in the zygopleura, to be next discussed, and which
also resemble them in being divisible only by one plane (the sagittal
median plane) into two equal halves. They differ, however, the amphipleural
body not being made up of two antimeres, but of at least three pairs of
antimeres (or three parameres), being therefore primitively radial. Hence
each of the symmetrical halves of the body contains more than one antimere.
Among the Radiolaria this form does not occur in the SPUMELLARIA,
ACANTHARIA, or PHÆODARIA; it is very common, however, among the
NASSELLARIA; many #Cyrtoidea# multiradiata and #Spyroidea# multiradiata
show this bilaterally radial ground-form, inasmuch as the body consists of
two symmetrical halves, and is also composed of numerous (usually three,
six, nine, or more) radial parameres. In the multiradiate Dicyrtida and
Tricyrtida the cephalis (the first joint) is usually bilateral, whilst the
thorax (the second joint) is multiradial. (See Gener. Morphol., Bd. i. pp.
495-506.)
38. _The Zygopleural Ground-Forms._--As zygopleural or dipleural
ground-forms, as opposed to the amphipleural, are classed those zeugites or
centroplana which are known {xviii}as "bilaterally symmetrical" in the
strictest sense of the term. This is the most important ground-form in the
animal kingdom, inasmuch as it obtains almost exclusively among the higher
animals (Vertebrata, Articulata, Mollusca, Vermes). The body consists of
only two antimeres, which correspond to the two symmetrical halves of the
body. Of the three dimensive axes two are allopolar, one isopolar; the oral
pole of the longitudinal main axis is different from the aboral; the dorsal
pole of the sagittal axis is different from the ventral; but the right pole
of the frontal axis is equal to the left. The right antimere is usually
precisely similar to the left (Eudipleura), more rarely it is slightly
dissimilar or asymmetrical (Dysdipleura). Among the Radiolaria this
ground-form is entirely wanting in the Porulosa or Holotrypasta
(SPUMELLARIA and ACANTHARIA), but on the contrary it is very common in the
Osculosa or Merotrypasta (NASSELLARIA and PHÆODARIA). In the NASSELLARIA it
is of special importance, for the typical _Cortina_ (the combination of the
primary sagittal ring with the basal tripod) exhibits the zygopleural
ground-form clearly sketched out; indeed it is usually clearly seen even in
the sagittal ring itself, for its ventral segment is more strongly curved
than the dorsal; its basal (or oral) pole is always different from the
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