Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII — John Shaqi
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
The important signification of the triradial structure, recurring in the
most different groups of NASSELLARIA, has been already pointed out
sufficiently by myself and by R. Hertwig. But the triradial #Plectoidea#
offer also another interesting relation of this characteristic structure,
some simple forms of this order appearing nearly identical with the
isolated triradial spicula of certain #Beloidea# (Thalassosphærida and
Sphærozoida). Even some more complex quadriradial and sexradial forms of
the latter reappear in exactly the same shape also in the former. This
identity may be perhaps an important indication of true affinity (compare
below).
The simplest and probably the most original kind of triradial structure is
exhibited by the genera _Triplagia_ and _Triplecta_ (Pl. 91, figs. 2, 7).
Here three equal radial spines lie in one horizontal plane and are united
in a common central point at equal angles, so that three lines connecting
their distal ends form a regular equilateral triangle. Simple triradial
spicula of the same regular form are also found in many {901}#Beloidea#
(_Lampoxanthium_, _Sphærozoum_, &c., Pls. 2 and 4). The central capsule of
these simplest #Plectoidea# (with vertical main axis) rests perpendicularly
on the horizontal triangle, formed by the triradial skeleton; the porochora
of the former (or the "area porosa") rests upon the central point of the
latter.
Another kind of triradial structure characterises the genera _Plagiacantha_
and _Plectophora_. The three radial spines united in the central point lie
here not in one plane, but diverge in different planes, so that they
correspond to the three lateral edges of a three-sided pyramid. Commonly
the three spines are of equal size, and also the angles between them equal,
so that the pyramid is regular, sometimes very flat, at other times more
elevated. Spicula of exactly the same form are also found in some
#Beloidea#. Probably the three divergent spines are homologous to the
three basal feet of numerous #Spyroidea# and #Cyrtoidea#. The central
capsule, according to Hertwig, is placed in the apical part of the pyramid,
the axes of both being identical, and the porochora resting in the apex
itself. This fact seems to contradict the above-mentioned affinity; but
since in _Triplecta_ and _Triplagia_ the three spines lie horizontally,
they may have changed this original position in different direction, in
_Plagiacantha_ and _Plectophora_ becoming divergent upwards, whereas in
_Plagoniscus_ and _Plectaniscus_ (as in the #Spyroidea# and #Cyrtoidea#)
directed downwards.
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