Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
The physiological value of the skeleton, with regard to the central
capsule, is different in the Plagonida and Plectanida; in the former it
supports, in the latter it encloses the capsule like a shell. In the
Nassellida, where no skeleton is developed, the central capsule is quite
free and naked, enveloped only by the calymma.
{906}The calymma or the extracapsular jelly-veil in all #Plectoidea# is
voluminous, and encloses not only the central capsule completely, but also
the skeleton wholly or partially. Its form is of the greatest value for the
development and configuration of the skeleton. Sometimes the calymma is
alveolate and foamy, as in _Nassella_ and the common _Thalassicolla_. In
several other #Plectoidea# the calymma seems to include numerous small
vacuoles, sometimes also pigment-granules. Xanthellæ are commonly scattered
in it in great numbers. The pseudopodia, arising in a large bunch from the
porochora of the capsule, and running along the branches of the radial
spines, seem to be always numerous, richly branched, and with a strong
tendency to form anastomoses. The peculiar form of their network is often
exactly preserved in the conformation of the skeleton, produced by them.
The peculiarities of this network require further accurate observations, as
does the whole organisation of the #Plectoidea#.
_Synopsis of the Families of_ #Plectoidea#.
I. Skeleton (originally tripodal) composed of radial
spines united in a common centre and supporting the
central capsule, without wicker-work, 1. PLAGONIDA.
II. Skeleton (originally tripodal) wattled, with irregular
wicker-work, composed of the united branches of radial
spines and enveloping the central capsule, 2. PLECTANIDA.
Family XLVI. #PLAGONIDA#, Haeckel.
_Plagonida_, Haeckel, 1881, Prodromus, p. 423.
_Plagiacanthida_ (_sensu strictiori_), Richard Hertwig, 1879, Organismus
d. Radiol., p. 72.
_Definition._--#Plectoidea# with a spiny skeleton, composed of radial
spines, which arise from a common central point or central rod, and support
the free central capsule.
The family #Plagonida# comprises those NASSELLARIA in which the skeleton is
only composed of united radial spines, arising from a common centre,
without any connection of meeting branches of the spines; the rudimentary
skeleton exhibits therefore neither a loose wickerwork (as in the closely
allied Plectanida), nor a ring (as in the #Stephoidea#), nor a complete
lattice-shell (as in the #Cyrtellaria#, the #Spyroidea#, #Botryodea#, and
#Cyrtoidea#). The central capsule, which possesses all the characters of
the MONOPYLEA, is therefore free, not enclosed, and only on one side
supported or partly protected by the radial spines or their branches.
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