Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
A. The number of joints into which the shell is divided by transverse
constrictions, serves here for the distinction of four primary groups or
suborders of the #Cyrtoidea#, viz., (1) Monocyrtida with one joint; (2)
Dicyrtida with two joints; (3) Tricyrtida with three joints; and (4)
Stichocyrtida with four or more joints. In my Prodromus (1881, p. 426) I
divided the latter group into Tetracyrtida (with four joints), and
Stichocyrtida (with five or more joints); but these two groups may be
united, since the fourth and all the succeeding joints are of rather
indifferent shape and of little morphological value. The three first
joints, however, are usually very different and possess a high
morphological importance, so that we distinguish the first joint as
cephalis, the second as thorax, and the third as abdomen. The uppermost
transverse constriction, which separates the two first joints, cephalis and
thorax, is the collar stricture and is usually caused by an internal
fenestrated septum, the cortinar septum. The second constriction, which
separates the second and third joints (thorax and abdomen) is called the
lumbar constriction. The following constrictions (in the Stichocyrtida) are
indifferent and of little morphological interest, and require therefore no
peculiar designation.
{1129}B. The radial structure, indicated by radial apophyses arising from
the shell, offers three principal differences, according to which the whole
group of #Cyrtoidea# may be divided into three large groups or sections,
viz., (1) Pilocyrtida, or Cyrtoidea triradiata, with three radial
apophyses; (2) Astrocyrtida, or Cyrtoidea multiradiata, with numerous
radial apophyses (four to nine or more); and (3) Corocyrtida, or Cyrtoidea
eradiata, without external radial apophyses. The majority of #Cyrtoidea#
are Pilocyrtida, with three radial apophyses, which are probably homologous
to the three primary feet of the #Plectoidea# and of _Cortina_ (therefore
"cortinar feet"). The Astrocyrtida, or the #Cyrtoidea# with a variable
number of radial apophyses (at least four to six) may be derived from the
Pilocyrtida by interpolation of secondary or interradial apophyses between
the three primary or perradial apophyses. The Corocyrtida, however, or the
#Cyrtoidea# without external radial apophyses, may have originated by
reduction and loss of the latter, either from the Pilocyrtida or from the
Astrocyrtida.
C. The shape of the basal mouth in the #Cyrtoidea# exhibits two essential
differences only, viz., (1) the terminal mouth of the shell is a simple
wide opening in the Stomocyrtida, or (2) the terminal mouth is closed by a
lattice-plate, in the Clistocyrtida. As these two different cases occur in
all the twelve families, which we have distinguished according to the
differences in the number of joints and in the radial structure, we get
altogether twenty-four subfamilies which are synoptically arranged in the
following table:--
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